Dacnusa (Dacnusa) reducta, Papp, 2004
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publication ID |
https://doi.org/ 10.5281/zenodo.12586698 |
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persistent identifier |
https://treatment.plazi.org/id/03C6463D-FF97-FFF2-FD80-5BA1FB734A60 |
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treatment provided by |
Felipe |
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scientific name |
Dacnusa (Dacnusa) reducta |
| status |
sp. nov. |
Dacnusa (Dacnusa) reducta View in CoL sp. n. fm
( Figs 63–74 View Figs 63–79 )
Material examined (18 f + 9 m). – Female holotype (No. 10670) + two female paratypes (Nos
10671–10672): Mongolia, Uvs aimak, Sandgebiet Altan els, 35 km WNW von Somon Tes, 1400 m, NIXON: 59 = first tergite, 60 = propodeum, 61 = head in dorsal view, 62 = mandible
23 Juni 1968, leg. KASZAB (loc. no. 1007). – One male paratype (No. 10688): Mongolia, Ostgobi aimak, 25 km SO von Čojren, 1200 m , 30 Juni 1963, leg. KASZAB (loc. no. 68). – One male paratype (No. 10689): Mongolia, Südgobi aimak, Gurban Sajchan ul, 30 km S von Somon Bulgan, 1700 m , 19 Juni 1964, leg. KASZAB (loc. no. 154). – One male paratype (No. 10690): Mongolia, Chentej aimak, Čandagan tal, 40 km O von Somon Žargaltchan, 1300 m , 28 Juli 1965, leg. KASZAB (loc. no. 316). – One female (No. 10673) + one male paratype (No. 10691): Mongolia, Chentej aimak, 150 km ONO von Öndörchaan, 10 km S vom Kerulen , 30 Juli 1965, leg. KASZAB (loc. no. 331). – One male paratype (No. 10692): Mongolia, Suchebaator aimak, 44 km SSW von Baruun urt, 1050 m , 2–3 August 1965, leg. KASZAB (loc. no. 349). – Two female paratypes (Nos 10674–10675): Mongolia, Suchebaator aimak, 15 km (1 f) and 45 km (1 f) N von Somon Erdenzagan, 950 m (1 f) and 900 m (1 f), 9 August 1965, leg. KASZAB (loc. no. 380 and 381). – One female paratype (No. 10676): Mongolia, Čojbalsan aimak, 50 km SO von Čojbalsan, 700 m , 16 August 1965, leg. KASZAB (loc. no. 421). – One male paratype (No. 10693): Mongolia, Central aimak, cca 30 km O von Somon Nalajch, 1530 m , 14 Juni 1966, leg. KASZAB (loc. no. 523). – One male paratype (No. 10694): Mongolia, Central aimak, 13 km W von Somon Bajanzogt, 1450 m , 17 Juni 1966, leg. KASZAB (loc. no. 527). – One female paratype ( No. 10677): Mongolia, Bulgan aimak, cca 20 km W von Somon Bajanuur, 1100 m , 18 Juni 1966, leg. KASZAB (loc. no. 531). – One female paratype (No. 10678): Mongolia, Central aimak, Ulan Baator , Nucht im Bogdo ul, 1650 m , 3 Juni 1967, leg. KASZAB (loc. no. 762). – Four female paratypes (Nos 10679–10682): Mongolia, Mittelgobi aimak, Choot bulag, zwischen Chuld und Delgerchangaj, 1480 m , 10 Juni (3 f) and 13 Juni (1 f) 1967, leg. KASZAB (loc. nos 782: 3 f, 913: 1 f). – Two female (Nos 10683–10684) and two male (Nos 10695–10696) paratypes: Mongolia, Central aimak, 12 km S von Somon Bajanbaraat, 1380 m , 13 Juni 1967, leg. KASZAB (loc. no. 918). – One female paratype ( No. 10685): Mongolia, Bulgan aimak, 30 km NNW von Somon Daschinčilen, 1200 m , 15 Juni 1968, leg. KASZAB (loc. no. 959). – One female paratype (No. 10686): Mongolia, Bulgan aimak, Namnan ul Gebirge , 23 km NW von Somon Chutag, 1150 m , 17 Juni 1968, leg. KASZAB (loc. no. 973). – One female paratype ( No. 10687): Mongolia, Chövsgöl aimak, 3 km SW von Somon Burenchaan, 1650 m , 21 Juni – 16 Juli 1968, leg. KASZAB (loc. no. 993).
Female holotype and 27 paratypes (18 f + 9 m) are deposited in the Hungarian Natural History Museum ( Department of Zoology ), Budapest, Hym. Typ. Nos 10670 (holotype), 10671–10687 (female paratypes) and 10688–10696 (male paratypes) .
Holotype is in good condition: glued on a pointed card by the right side of the meso- and metasoma. Paratypes are also in good condition: glued on pointed cards either by their sides or by mesosternum, three specimens with damaged flagelli, two specimens with creased wings.
Etymology. – The name of the new species “reducta ” refers to the conspicuously shortened marginal cell of the fore wing.
Description of the female holotype. – Body 1.5 mm long. Antenna shorter than body and with 14 antenomeres. First three flagellomeres equal in length (four times longer than broad apically), further flagellomeres gradually shortening and thickening so that penultimate flagellomere 1.6 times as long as broad. – Head in dorsal view ( Fig. 63 View Figs 63–79 ) less transverse, 1.7 times as broad as long, temple beyond eye swollen, i.e. head between temples broader than between eyes; temple 1.3 times as long as eye. Ocelli small, OOL one-fourth longer than POL. Eye in lateral view almost twice as high as wide, temple almost twice as wide as eye. Mandible moderately broadening distally, third tooth reduced ( Fig. 64 View Figs 63–79 ). Head polished.
Mesosoma in lateral view 1.25 times as long as high. Notaulix entirely indistinct. Mesoscutum bare and polished, its anterior declivous part punctured and haired. Precoxal furrow distinct as short and very shallow impression. Propodeum smooth, shiny, laterally punctured and hairy. – Hind femur four times as long as broad medially ( Fig. 65 View Figs 63–79 ). Hind tibia and tarsus equal in length.
Fore wing about as long as body. Pterostigma ( Fig. 66 View Figs 63–79 ) 6.6 times as long as wide, almost parallel-sided, issuing r from its proximal fourth, r half as long as width of pterostigma; marginal cell narrow and short, i.e. 3–SR + SR1 bent and ending far before tip of wing; 1–R1 very short, 0.25 times as long as length of pterostigma; m–cu antefurcal ( Fig. 67 View Figs 63–79 , see arrow); first subdiscal cell as in Fig. 68 View Figs 63–79 , CU1b depigmented.
First tergite short and distinctly broadening posteriorly, somewhat wider behind than long, pair of spiracles just before middle of tergite, scutum smooth and shiny, tergite laterally from scutum uneven ( Fig. 69 View Figs 63–79 ). Second and third tergites fused, i.e. border between them absent, together with further tergites polished. Ovipositor sheath short, in lateral view as long as third tarsomere of hind tarsus ( Fig. 70 View Figs 63–79 ).
Scape brown, pedicel light brown, flagellum dark brown. Head and mesosoma dark brown, metasoma brown. Mandible and palpi yellowish brown. Tegula dark brown, parategula light brown. Legs brown, fore femur apically and fore tibia basally yellowish. Wings hyaline, pterostigma and veins brownish yellow.
wing. – 79. Dacnusa (Dacnusa) pseudolugens TOBIAS : antefurcal m–cu (↓)
Description of the 17 female paratypes. – Similar to the female holotype. Body 1.2–1.5 mm long (1.2: 3 f, 1.3: 11 f, 1.4: 2 f, 1.5: 1 f). Antenna with 13–15(–18) antennomeres (13: 6 f, 14: 4 f, 15: 6 f, 18: 1 f). Head in dorsal view 1.65–1.7(–1.75) times as broad as long. Vein m–cu either (just) antefurcal ( Fig. 67 View Figs 63–79 ) or interstitial ( Fig. 71 View Figs 63–79 ). Hind femur 3.8–4 times as long as broad somewhat distally. First tergite usually somewhat broader behind than long ( Fig. 69 View Figs 63–79 ) or, less usually, as long as broad behind (3 f, Fig. 72 View Figs 63–79 ). Ground colour of body blackish with faint brownish tint.
Description of the nine male paratypes. – Similar to the female types. Body 1.2–1.5 mm long (1.2: 4 m, 1.3: 3 m, 1.4: 1 m, 1.5: 1 m). Antenna with 14–17 antennomeres (14: 3 m, 15: 3 m, 16: 1 m, 17: 2 m). Head in dorsal view either similar to the that of female ( Fig. 63 View Figs 63–79 ) or temple somewhat less swollen ( Fig. 73 View Figs 63–79 ). First tergite less broadening posteriorly, as long as broad behind ( Fig. 74 View Figs 63–79 ). Ground colour of body blackish.
Distribution: Mongolia. – Host unknown.
The new species, Dacnusa (Dacnusa) reducta , is nearest to D. (D.) lugens (HALIDAY, 1839) considering their tiny body and reduced length of marginal cell; the two species are distinguished by the features keyed:
1 (2) Marginal cell short, 1–R1 half (0.5) as long as pterostigma ( Fig. 75 View Figs 63–79 ); m–cu always antefurcal (cf. Figs 67 and 79 View Figs 63–79 , see arrows). Temple in dorsal view not swollen, i.e. head between temples as broad as between eyes, eye and temple of equal length ( Fig. 76 View Figs 63–79 ). Antenna with 17–23 antennomeres (fm). Mandible with distinct third tooth and less broadening distally ( Fig. 77 View Figs 63–79 ). f: 1.5–1.7 mm, m: 1.5–1.6 mm D. (D.) lugens (HALIDAY)
2 (1) Marginal cell very short, 1–R1 0.3 times as long as pterostigma ( Fig. 66 View Figs 63–79 ); m–cu either (just) antefurcal ( Fig. 67 View Figs 63–79 , see arrow) or interstitial ( Fig. 71 View Figs 63–79 , see arrow). Temple in dorsal view swollen, i.e. head between temples broader than between eyes, temple clearly longer than eye ( Figs 63, 73 View Figs 63–79 ). Antenna with 13–15 (f) and 17 (m) antennomeres. Mandible with reduced third tooth and more broadening distally ( Fig. 64 View Figs 63–79 ). f: 1.3–1.5 mm, m: 1.5 mm
D. (D.) reducta sp. n. Dacnusa (Pachysema) aterrima THOMSON, 1895 is similar to D. (D.) reducta sp. n. considering their dark coloured body (legs too) and shortened marginal cell, their distinction is presented as follows:
1 (2) First tergite long ( Fig. 80 View Figs 80–91 ), (1.2–)1.3–1.4 times as long as broad behind (subgeneric feature). Marginal cell of fore wing less narrow ( Fig. 78 View Figs 63–79 ). Antenna with 16–19 antennomeres (fm). Hind tibia somewhat longer than hind tarsus. fm: 1.5–2 mm D. (P.) aterrima THOMSON View in CoL
2 (1) First tergite short (subgeneric feature), usually broader behind than long ( Fig. 69 View Figs 63–79 ) or, in male and less usually in female, as long as broad behind ( Figs 72, 74 View Figs 63–79 ). Marginal cell of fore wing narrow ( Fig. 66 View Figs 63–79 ). Antenna with 13–15 (f) and
17 (m) antennomeres. Hind tibia and tarsus equal in length. f: 1.3–1.5 mm, m: 1.5 mm D. (D.) reducta View in CoL sp. n.
With the help of TOBIAS’ s key (1998: 324–330) to the Dacnusa species of Russian Far East the new species, Dacnusa (D.) reducta , runs to D. pseudolugens TOBIAS, 1998 (Asiatic Russia: Magadan Region, Maritime Territory, Yakutia), the two species are separated by a few features as follows:
1 (2) Antenna with 21 antennomeres. Fovea of mesoscutum present. Pterostigma 2–2.5 times as long as 1–R1, m–cu clearly antefurcal ( Fig. 79 View Figs 63–79 ). Temple in dorsal view either as long as (m, Fig. 81 View Figs 80–91 ) or slightly longer than eye (f, Fig. 82 View Figs 80–91 ), weakly swollen beyond eye. Third tooth of mandible less reduced (cf. Fig. 77 View Figs 63–79 ). f: 77). f: 1.5–1.7 mm D. (D.) pseudolugens TOBIAS View in CoL
2 (1) Antenna with 13–15(–18) antennomeres. Fovea of mesoscutum absent. Pterostigma four times as long as 1–R1, m–cu either antefurcal ( Fig. 67 View Figs 63–79 , see arrow) or interstitial ( Fig. 71 View Figs 63–79 , see arrow). Temple in dorsal view 1.3–1.4 times as long as eye, clearly swollen beyond eye ( Fig. 63, 73 View Figs 63–79 ). Third tooth of mandible reduced ( Fig. 64 View Figs 63–79 ). fm: 1.3–1.5 mm D. (D.) reducta View in CoL sp. n.
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