Gromphas lacordairii (Oken, 1834)

Cupello, Mario, 2024, The genus Gromphas Dejean, 1836 (Coleoptera, Scarabaeinae): nomenclature, distribution, and conservation, including a contribution to the debate on electronic publications in zoology, Zoosystema 46 (2), pp. 23-59 : 52-53

Gromphas lacordairii :

Argentina. Autonomous City of Buenos Aires • 1 unsexed specimen; Palermo; IV.1943; no collector; ex Martínez collection; CMNC. — Buenos Aires • 1 ♂; Boulogne Sur Mer; XI.1941; no collector; ex Martínez collection; CMNC • 1 unsexed specimen; Florencio Varela; I.1978; Giacomozzi leg.; CMNC • 1 unsexed specimen; San Isidro ; 14.IV.1956; H. J. Molinari leg; CMNC • 1 unsexed specimen; San Isidro; XII.1967; no collector; ex Martínez collection; CMNC • 1 unsexed specimen; San Isidro ; XI.1973; no collector; ex Martínez collection; CMNC. — Chaco • 1 ♂, 3 ♀; Presidencia de la Plaza; 26°59’S, 59°50’W; 7.X.2006; M. Damborsky leg.; TAMU. — Corrientes • 1 unsexed specimen; ‘ 5 km W Ituzaingó’; IX.1979; no collector; ex Martínez collection; CMNC • 2 unsexed specimens; Ituzaingó; III.1976; no collector; ex Martínez collection; CMNC. — Entre Ríos • 16 ♂, 7 ♀; Uruguay, Molino, Pronunciamiento; I-III.1963; J. Foerster leg.; cow dung; TAMU. — Misiones • 1 ♀; Cainguás, Dos de Mayo; I.1966; J. Foerster leg.; carrion; TAMU. — Salta • 1 unsexed specimen; Embalse de Cabra Corral; 18.I.1984; Bordón leg.; ex Martínez collection; CMNC • Guachipas; I.1948; no collector; ex Martínez collection; CMNC • 1 unsexed specimen; Guachipas, Alemanía; III.1973; no collector; ex Martínez collection; CMNC. — Santiago del Estero • 2 unsexed specimens; Río Hondo, Termas de Río Hondo ; 16.II.1982; Henry Howden and Anne Howden leg.; CMNC. — Tucumán • 1♂; Yerba Buena, San Javier , 1200 m; 20.I.1970; Mielke leg.; DZUP .

Bolivia. Santa Cruz • 1 unsexed specimen; Chiquitos, Roboré, Santiago de Chiquitos ; XI.1959; no collector; ex Martínez collection; CMNC . — Tarija • 1♀; between Yaguacua and Caiza; 21°50’52”S, 63°36’26”W; 620 m; 3.I.2005; Mann, Hamel and Herzog leg.; horse dung; Chaco transition zone, low lying pasture, open habitat, sandy-loam; TAMU .

Brazil. Espírito Santo • 4 ♂, 1 ♀; Linhares ; XI.1965; A. Maller leg.; DZUP • 2 ♂; Linhares ; XII.1965; A. Maller leg.; DZUP . — Mato Grosso • 1 ♀; Santo Antonio de Leverger ; XII.1965; W. D. Edmonds leg.; meat trap; ‘capoeira’ vegetation; TAMU • 4 ♂ [1 with genitalia dissected], 1 ♀; Santo Antonio de Leverger ; 4.XII.1965; W. D. Edmonds leg.; cattle dung; TAMU . — Mato Grosso do Sul • 1 ♂; Corumbá ; 17.X.1961; F. M. Oliveira leg.; DZUP . — Paraná • 1♂, 1 ♀; Araucária, Barigui ; V.1941; no collector; ex F. Justus Júnior collection; DZUP • 1 ♀; Curitiba ; X.1961; S. Laroca leg.; DZUP • 5 ♂, 7 ♀; Curitiba ; 25.I.1962; S. Laroca leg.; DZUP • 5 ♂, 5 ♀; Curitiba ; 28.X.1964; Sebastião Laroca leg; DZUP • 1♂, 1 ♀; Curitiba ; 21.XI.1965; Mitchell and Graf leg.; DZUP • 1 ♂; Curitiba ; 11.IV.1968; Gonçalves leg.; DZUP • 1 ♀; Curitiba ; 900 m; 2.XI.1963; Departamento de Zoologia da Universidade Federal do Paraná leg. (‘ Dept. ZOO. leg’); DZUP • 1 unsexed specimen; Curitiba, Cajuru ; X.1938; Claretiano leg; MGAP • 1 unsexed specimen; Curitiba, Parolin (‘ Parolim’ ); I.1937; Claretiano leg; MGAP • 1 ♀; Guaíra ; X.1953; C. R. Leite leg.; ex Alvarenga collection; DZUP • 1 ♀; Ponta Grossa ; VIII.1942; no collector; ex F. Justus Júnior collection; DZUP • 1 ♂, 3 ♀; São José dos Pinhais ; 18.XI.1966; Laroca and Antoni leg.; DZUP. — Rio de Janeiro • 1 ♀; Duque de Caxias, São Bento (‘ S Bento’); XII.1954; P. A. Tales leg.; ex Alvarenga collection; DZUP • 1 ♀; Rio de Janeiro, Botafogo ; X.1957; Alvarenga leg.; ex Alvarenga collection; DZUP • 1 unsexed specimen; Rio de Janeiro, Copacabana ; XII.1994; R. L. Vaz de Mello leg.; CMNC . — Rio Grande do Sul • 3 ♂, 1 ♀; Cachoeira do Sul ; II.1996; H. Schaurich leg.; MCNZ • 1 ♂; Capão da Canoa, Curumim ; 22.I.1978; C. J. Becker leg.; MCNZ • 1 ♀; Capão da Canoa ; 9.X.1989; M. A. Santos leg.; MCNZ • 3♂, 4 ♀; Capão da Canoa ; 18.XI.1989; M. A. Santos leg.; MCNZ • 1 unsexed specimen; Cerro Largo ; I.1931; no collector; MGAP • 1 unsexed specimen; Cerro Largo ; I.1939; no collector; MGAP • 1 unsexed specimen; Cerro Largo ; I.1940; no collector; MGAP • 2 unsexed specimens; Cerro Largo ; 1949; no collector; MGAP • 2 unsexed specimens; Cerro Largo ; I.1949; no collector; MGAP • 12♂, 14 ♀; Charqueadas, Fazenda Águas Claras ; 12.I.1989; A. Ferreira leg.; MCNZ • 2 ♂, 2♀; Cidreira, Salinas ; 2.XI.1992; Lascombe leg.; MCNZ • 1 ♂; General Câmara (‘ Gen Câmara’ ); 16.IX.1982; A. Lise leg.; MCNZ • 4 ♂, 5♀; Imbé ; II.1961; Buckup leg.; MCNZ • 1♂, 1 ♀; Imbé, Santa Teresinha Beach ; II.1973; M. H. Galileo leg.; MCNZ ; • 1♂; Imbé, Santa Teresinha Beach ; 6.XI.1999; L. Moura and I. Heydrich leg.; MCNZ • 3 unsexed specimens; Mostardas ; I.1945; no collector; MGAP • 16 unsexed specimens; Nova Petrópolis ; I.1928; no collector; MGAP • 5 unsexed specimens; Osório ; 3.I.1950; no collector; MGAP • 8 unsexed specimens; Osório ; 24.I.1958; no collector; MGAP • 1 unsexed specimen; Osório ; 24.I.1958; Pio Buck leg.; MGAP • 1 unsexed specimen; Osório ; 25.I.1958; no collector; MGAP • 1 unsexed specimen; Osório ; 25.I.1958; Pio Buck leg.; MGAP • 1♀; Osório ; 2.II.1964; P. Braun leg.; MCNZ • 1 unsexed specimen; Pareci Novo ; 20.X.1927; no collector; MGAP • 1 unsexed specimen; Pareci Novo ; VIII.1935; no collector; MGAP • 1 unsexed specimen; Pareci Novo ; XI.1938; no collector; MGAP • 1 unsexed specimen; Pelotas ; II.1963; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 16.X.1928; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 29.IX.1929; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 29. V.1934; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 15.XI.1935; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 1941; ‘ T. L.’ leg.; MGAP ; • 1 unsexed specimen; Porto Alegre ; 1943; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 25.IX.1949; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 3.X.1956; no collector; MGAP • 1 ♀; Porto Alegre ; 6.XII.1962; P. Braun leg.; MCNZ • 1 unsexed specimen; Porto Alegre ; 7.X.1963; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 22.X.1963; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 1964; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 26.I.1965; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 7.II.1965; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 7.X.1965; no collector; MGAP • 1 unsexed specimen; Porto Alegre ; 26.X.1966; no collector; MGAP • 1 ♀; Porto Alegre ; 18.VIII.1970; no collector; MCNZ • 1 ♂; Porto Alegre ; 8.XI.1993; V. L. C. Lascombe leg.; MCNZ • 3 unsexed specimens; Porto Alegre, Belém Novo ; I.1959; no collector; MGAP • 1 unsexed specimen; Quaraí ; I.1945; no collector; MGAP • 1 unsexed specimen; Santo Augusto ; 12.X.1966; O. Roppa leg.; CMNC • 2 ♂; São Jerônimo, Fazenda Santa Hermenêutica ; 13.I.1989; A. Ferreira leg.; MCNZ • 3 ♂, 3♀; São Jerônimo, Instituto Rio Grandense do Arroz (‘IRGA’); 11.I.1989; on dung; A. Ferreira leg.; MCNZ • 1 unsexed specimen; São Leopoldo ; XI.1958; no collector; MGAP • 1♂; Torres ; X.1956; T. de Lema leg.; MCZN • 1 ♂; Triunfo, Braskem plant (stated on the label by its former name ‘COPESUL’); 21.IX.1989; M. Hoffmann leg.; MCNZ • 1 ♀; Triunfo, Braskem plant (‘COPESUL’); 14-15.I.1997; A. Franceschini leg.; MCNZ • 2 ♀; Uruguaiana, Estância Corumbá ; 18.XII.1996; N. S. Macedo leg.; MCNZ • 1 ♂, 1♀; Viamão ; X.1956; L. Buckup and E. Buckup leg.; MCNZ • 1 ♀; Xangri-Lá, Atlântida Beach ; 20.I.1968; no collector; MCNZ . — Santa Catarina • 1 unsexed specimen; ‘ Emboebas’ (?); 3.IV.1953; no collector; MGAP • 1 ♀; Canoinhas, Pinhal; XII.1952; A. Maller leg.; ex Campos Seabra collection; TAMU • 1 unsexed specimen; Itapiranga ; IX.1953; MGPA • 1 unsexed specimen; Itapiranga ; V.1954; no collector; MGAP • 1 unsexed specimen; Joinville ; VI.1949; no collector; MGAP • 1 unsexed specimen; São Bento do Sul; I.1989; no collector; CMNC • 1 unsexed specimen; Seara, Nova Teutônia ; 25.IX.1963; F. Plaumann leg.; CMNC • 1 unsexed specimen; Seara, Nova Teutônia ; 23.X.1963; F. Plaumann leg; CMNC • 1 unsexed specimen; Seara Nova Teutônia ; 2.I.1964; F. Plaumann leg.; CMNC • 1 unsexed specimen; Seara, Nova Teutônia ; 30.I.1964; F. Plaumann leg.; CMNC • 6 unsexed specimens; Seara, Nova Teutônia ; XII.1970; no collector; CMNC • 1 unsexed specimen; Seara, Nova Teutônia ; 300-500 m; II.1970; no collector; CMNC • 1♂; Seara, Pinhal , 700 m; IV.1958; no collector; DZUP • 1 ♂; Seara, Pinhal , 700 m; XII.1958; no collector; DZUP . — São Paulo • 1 unsexed specimen; ‘CIPO’ (?); 6.II.1973; V. N. Alin leg.; CMNC • 1 unsexed specimen; ‘CIPO’ (?); 12.XII.1973; V. N. Alin leg.; CMNC • São Paulo, Santo Amaro (‘ S. Amaro’); 20.XII.1953; no collector; MGAP .

Paraguay. Alto Paraguay • 1♀; Puerto la Esperanza ; 2.XII.2006; C. Aguilar Julio leg.; CAPC . — Caaguazú • 1 unsexed specimen; Caaguazú ; XII.1948; F. H. Schade leg.; CMNC . — Caazapá • 1 unsexed specimen; Caazapá; 1-8.XII.1990; G. Arriagada leg.; CMNC. — Capital District • 1 ♂; Asunción ; 26.II.1992; J. A. Kochalka leg.; MNHNP • 1 unsexed specimen; Asunción, Mburicaó ; 12.X.1990; G. Arriagada leg.; CMNC • 1 unsexed specimen; Asunción, Mburicaó ; 7.I.1991; G. Arriagada leg.; CMNC. — Central • 1 ♂; Ypacaraí (‘ 1000 Mts. al E. de / YPACARAÍ ’); 23.X.1996; D. Moyano leg.; CAPC • 1 ♂; Ypacaraí (‘ 1000 Mts. al E. de / YPACARAÍ ’); 23.X.1996; D. Moyano leg.; MNHNP . — Concepción • 1♀; San Lázaro, Vallemí ; 26.XI.2006; C. Aguilar leg.; CAPC . — Guairá • 1 ♂; Cordillera del Ybytyruzú, Carpa Cué ; 24.XII.1994; Aguilar leg.; MNHNP • 2♀; Cordillera del Ybytyruzú, Cerro Akatí ; 19.II.2004; no collector; CAPC • 1 unsexed specimen; Mbocayaty (‘ Bcayaty’ ); 23.IX.1946; Schade leg.; CMNC . • 1 unsexed specimen; Villarrica ; X.1941; Schade leg.; CMNC . — Itapúa • 1 ♀; eastern portion of Isla Yacyretá, Paraná River [Butia savanna]; 14.XII.1999; J. A. Kochalka leg.; MNHNP . — Paraguarí • 1 unsexed specimen; Parque Nacional Ybycuí ; 13.I.1996; C. Aguilar J. leg.; CMNC . — San Pedro • 2 unsexed specimens; Cororo , Río Ypané ; II.1979; no collector; Martínez leg.; CMNC • 2♂; Nueva Germania ; 15- 20.I.1992; Aguilar leg.; MNHNP .

Uruguay. Canelones • 1 ♂; Pando ; 10.I.2005; no collector; CAPC • 6 unsexed specimens; Ruta 82, Km 46 500, Campo Experimental Instituto de Higiene ; 10.XI.2020; unknown collector; cow dung; FCUR . — Cerro Largo • 1 ♀; no further locality or date; Sáenz leg.; TAMU . — Montevideo • 1 unsexed specimen; no further data; MGAP . — Tacuarembó • 1 unsexed specimen; Ruta 43, on the banks of Río Negro near San Gregorio de Polanco ; 32°38’43”S, 55°47’45”W; 20.X.2019; unknown collector; cow dung; FCUR .

Likely erroneous data: Bolivia. Santa Cruz • 1 ♀; Ichilo, Buena Vista ; 380 m; XI.1968; F. Steinbach leg.; TAMU .

Unknown wild origin • 1 ♂; reared from a laboratory colony at Texas A&M University, College Station , TX, USA; summer 1981; ex G. T. Fincher collection; TAMU • 1 ♂; same data as for preceding but the date; XI.1981; ex G. T. Fincher collection; TAMU; 1♀ • same data as for preceding but the date; 1982; ex G. T. Fincher collection; TAMU .

No data • 2 ♂, 8 ♀; DZUP • 12 unsexed specimens; MGAP.