Panzeria bisetosa, Shima, 2020

Shima, Hiroshi, 2020, The genus Panzeria Robineau-Desvoidy (Diptera: Tachinidae) from Japan, Zootaxa 4816 (4), pp. 541-575 : 549-551

Panzeria bisetosa sp. nov.

( Figs 15–20 , 71–72 )

Diagnosis. This species is characteristic among species of Panzeria in having male terminalia with narrow cerci and surstylus and female 8th abdominal sternite absent. A rather small species; abdomen long ovate; 2 katepisternal setae; 4 postsutural dorsocentral setae; 2–3 presutural acrostichal setae; apical scutellar setae absent; wing hyaline, without tinge along r-m crossvein; male cerci narrow and curved dorsally.

Description. Body length: 6.3–8.7 mm.

Male. Head black in ground color, reddish brown on narrow triangular portion below eye and pale yellowish brown on lower portion of face; parafrontal, parafacial, gena and occiput with dense grayish-white pruinosity, somewhat silvery and more whitish on parafacial; face with whitish pruinosity; frontal vitta velvety dark brown; antenna dark brown, apical portion of pedicel reddish brown; palpus reddish yellow, narrowly darkened on basal portion. Frons at narrowest point about 0.22 of head width; frontal vitta weakly widened anteriorly, nearly as wide as parafrontal at middle; frons in lateral view about 1.3 times as long as face; parafacial 0.8–0.9 times as wide as postpedicel at mid height in horizontal position; gena about 1/2 of eye height in horizontal position (3/ 8 in profile); face weakly concave, lower margin warped forward, slightly expanded beyond base of vibrissa. Inner vertical seta rather fine, 1/2–5/8 as long as eye height; 9–11 frontal setae, lowest seta nearly level with apex of pedicel; ocellar setae about 3/4 as long as inner vertical seta; parafrontal with 2–3 rows of fine short black hairs, only 1–2 descending just below lowest frontal seta; parafacial bare; occiput bulged, with 2–3 rows of black hairs behind postocular setae. Antenna falling short of lower margin of face by about 1/4 length of pedicel; postpedicel about 2 times as long as wide, about 2 times as long as pedicel; arista with 2nd aristomere about 2 times as long as wide, 3rd aristomere thickened on its basal 2/5. Prementum short, about as long as postpedicel; palpus nearly cylindrical, only slightly widened near apex, about 4/5 as long as pedicel and postpedicel combined.

Thorax black in ground color, apex of scutellum sometimes narrowly reddish brown; scutum with rather thin grayish pruinosity, somewhat brownish near median portion; pleura with thin grayish pruinosity; scutum with 5 broad longitudinal vittae, median vitta sometimes indistinct on presutural area, outer vitta distinctly wider than pruinose portion between inner and outer vittae on postsutural scutum. Two posthumeral setae; 2 or 3 presutural and 3 postsutural acrostichal setae; 4 postsutural dorsocentral setae; scutellum with a pair of discal and 3 or 4 pairs of marginal setae, lateral scutellar seta sometimes single, apical setae absent; all marginal scutellar setae very long, about 2 times as long and scutellum, if 2 lateral setae present, then anterior one fine; subapical scutellar setae close at bases and parallel to each other, distance between their bases about 1/2 that between basal and subapical setae; 2 katepisternal setae; anepimeral seta extending at most to posterior margin of upper calypter; anatergite with several fine short hairs. Wing hyaline, weakly tinged with pale brown throughout, without dark tinge on r-m crossvein; calypter whitish; second costal sector about 2/5 length of 3rd and 3rd sector subequal in length to 4th; vein M nearly rightangled at bend, sometimes with a very short appendage, length of M from dm-cu crossvein to bend about 1.5 times distance between bend and wing margin, and about 3/4 as long as dm-cu crossvein. Legs black, hind tibia sometimes dark reddish brown ventrally; fore tibia with 4–5 anterodorsal, 1–2 posterodorsal and 2–3 posterior setae; mid tibia with 3 anterodorsal, 2–3 posterodorsal and 1 ventral setae; hind tibia with a row anterodorsal setae of irregular length, 3–5 posterodorsal and 2 ventral setae; claws and pulvilli long, subequal in length to 5th tarsomere.

Abdomen black in ground color; dorsum with whitish pruinosity on anterior 1/2–1/4 of 3rd tergite and anterior 1/2–2/3 of 4th and 5th tergites, pruinosity thinner medially and faded posteriorly on each tergite, a broad mid-dorsal longitudinal vitta distinct on 3rd to 5th tergites; venter with thin grayish-white pruinosity throughout. Hairs on dorsum suberect, dense and rather long, becoming sparser posteriorly, on venter fine, short and dense; syntergite 1+2 without median marginal setae and with 2 lateral marginal setae; 3rd tergite with a pair of strong median discal and median marginal setae and 2 lateral marginal setae; 4th tergite with a pair of strong median discal and a row of marginal setae; 5th tergite with regular rows of strong discal and marginal setae; 5th sternite with posterior lobe occupying about posterior 1/3 of its length, inner posterior corner of posterior lobe roundly produced, with dense long hairs; median membranous portion rather narrow, T-shaped.

Terminalia: 6th tergite to epandrium with very thin grayish pruinosity; 6th tergite rather long, fused with syntergosternite 7+8 medially, about 1/2 as long as syntergosternite 7+8, with a row of rather strong marginal setae and 1–2 rows of fine hairs; epandrium large, with a short posterior invagination; cerci in dorsal view strongly narrowed from base to basal 2/5, then almost parallel-sided to apex, in lateral view well curved medially, apex narrowly truncated; surstylus narrow, nearly straight in lateral view, with 2 short and strong spines at apex; hypandrial arms broad; pregonite wide, posterior margin curved outwardly with several long and short hairs; postgonite narrow, weakly widened posteriorly; epiphallus absent; basiphallus rather long; distiphallus broad rectangular in lateral view.

Female. Differs from male as follows: antenna with scape and pedicel more reddish, especially on bases and apices; frons at narrowest point about 0.31 of head width; frons in profile about 1/2 times as long as face; parafacial width about 0.83 of postpedicel (0.67 in profile); head setae stronger, inner vertical seta more than 3/4 length of eye height; outer vertical seta developed, about 2/3 as long as inner vertical seta; 1 outwardly directed prevertical seta, about 4/5 as long as outer vertical seta; ocellar setae nearly as long as outer vertical seta; 2 subequally long proclinate orbital setae, subequal in length to outer vertical seta; 6–8 frontal setae; antennal pedicel with a long seta which is shorter than pedicel; postpedicel 1.6–1.8 time as long as wide and 1.6–1.8 times as long as pedicel; palpus clavate, slightly longer than postpedicel; legs with claws and pulvilli shorter than 5th tarsomere; abdominal hairs shorter and appressed on 3rd and 4th tergites. Terminalia: 6th tergite broadly divided into short and high hemitergites, with a row of strong setae marginally; 7th hemitergite subequal in length to 6th hemitergite, lower than the latter, with a row of rather sparse strong setae medially; 7th sternite weakly constricted posteromedially, ventromedian portion weakly membranous; 8th tergite absent; epiproct divided medially into 2 small sclerites, each with a strong seta.

Type material. Holotype male: [ Japan] Kyushu / Fukuoka / Mt. Kanayama / 15.v.1984 / H. Shima ( BLKU).

Paratypes (all in KUM): Japan, Honshu : 1 female, Aomori Pref., Mt. Bonju , 4.vi.1983, S. Fukushi ; 1 male, Nagano Pref., Shimashimatani , 24.v.1975, A. Nakanishi & J. Emoto (Malaise trap). Kyushu : 1 female, Fukuoka City, Aburayama , 21–28.iv.1983, H. Shima (yellow pan trap) ; 1 female, Kyushu, Fukuoka City, Mt. Kanayama , 1.vi.1984, H. Shima ; 1 male, Fukuoka Pref., Mt. Kusenbu , 29.v.2009, H. Shima ; 1 male, Kumamoto Pref., Izumimura, Nitao , 29.iv.1974, I. Ohtsuka .

Etymology. The specific name refers to the presence of only two katepisternal setae in this species, combining the Latin noun bi, two, with the noun seta, hair.

Distribution. Japan (Honshu, Kyushu).

Figures

FIGURES 15–20. Male and female terminalia of Panzeria bisetosa sp. nov. (paratypes, Nagano, Honshu, Japan & Yamanashi, Honshu, Japan, KUM). 15. Male 5th abdominal sternite in ventral view (hairs omitted on left side). 16. Epandrium, cerci and surstylus in lateral view (hairs omitted). 17. Same in dorsal view (hairs omitted). 18. Hypandrium, pregonite, postgonite and phallus in lateral view. 19. Female terminalia in lateral view. 20. Same in ventral view (hairs omitted). Scale bars: 0.2 mm for 15, 0.1 mm for 16–20.

FIGURES 69–84. Panzeria spp. 69–70. Panzeria atriventris sp. nov. (Mt Zao, Honshu, Japan, KUM). 71–72. Panzeria bisetosa sp. nov. (Fukuoka, Kyushu, Japan, KUM). 73–75. Panzeria emdeni (Mesnil) (Nagano, Japan, KUM). 76–77. Panzeria japonica sp. nov. (Tokyo, Honshu, Japan, KUM). 78–79. Panzeria meridionalis sp. nov. (Kagoshima, Kyushu, Japan, KUM). 80–81. Panzeria rufitibialis sp. nov. (Iriki Pass, Kyushu, Japan, KUM). 82–84. Panzeria tuberculata (Zimin) (Nagano, Japan, KUM). 69, 71, 73, 77, 79–80, 82. Dorsal habitus. 70, 72, 76, 78, 81. Lateral habitus. 74, 84. Male head in dorsal view. 75, 83. Male thoracic pleura (anterior part) in lateral view.