Lysmus Navás, 1910

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 52-53

Lysmus Navás, 1910

( Figs 34–35 )

(= Eosmylus Krüger, 1915: 73. Type species: Spilosmylus nigricornis Nakahara, 1914: 509 )

(= Neolysmus Nakahara, 1955: 13 . Type species: Neolysmus ogatai Nakahara, 1955: 13 )

Type species. Osmylus harmandinus Navás, 1910: 190 (by original designation).

Diagnosis. Three ocelli present; antennae less than FW length; wings ovoid, hyaline, FW sometimes with dark markings; FW costal area broadened basally, subcostal veinlets simple, not arranged sub-radially at base (sometimes basal 2–3 arranged divergently); RP stem short; 10–13 RP branches present; end-twigging of radial and medial veins along wing margin regular and one level; FW with stem of RP relatively short, RP1 arising proximal to M fork; RP in both wings with two distinct gradate series, subparallel in orientation, more basal crossveins semi irregular in arrangement, sometimes with third poorly defined gradate series; FW M vein fork near base of wing, between first and second branches of RP; FW CuP with more branches than CuA.

Comments. Lysmus was previously placed in Spilosmylinae , underscoring the difficulties in differentiating and classifying members of both subfamilies. Winterton et al. (2017) recovered this genus as the sister to Gryposmylus , and Winterton & Wang (2016) noted that the genera are differentiated by features such as the orientation of the basal subcostal veinlets and gradate series crossveins, and by the position of the M fork relative to the origin of RP1. Considering the variability of some venational features in Lysmus , this genus should be a priority for revision to clarify its status relative to other genera in the subfamily, especially genera such as Gryposmylus , Osmylidia and Protosmylus .

Included species.

L. beccarii ( Navás, 1929) ( Indonesia, Japan)

L. canimaculatus Jia & Liu, 2005 ( China)

L. harmandinus ( Navás, 1910) ( Japan, Russia) (= Lysmus kurilensis Kuwayama, 1956 ; = Spilosmylus nigricornis Nakahara, 1914 )

L. oberthurinus ( Navás, 1910) ( China)

L. ogatai ( Nakahara, 1955) ( Japan, China)

L. pallidius Yang & Liu, 2001 ( China)

L. qingyuanus Yang et al., 1995 ( China)

L. victus Yang, 1997 ( China)

L. zanganus Yang et al. in Huang et al., 1988 ( China)

Figures

FIGURE 34. Protosmylinae. Lysmus harmandinus Navás.

FIGURE 35. Protosmylinae. Lysmus victus Yang.