Kempynosmylus Makarkin, 2014

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 29

Kempynosmylus Makarkin, 2014

( Fig. 11C )

Type species. Kempynosmylus zherikhini Makarkin, 2014: 9 (by original designation).

Diagnosis. Forewing ovoid; RP with numerous branches (>16), strongly sinuate distally; numerous regularly arranged crossveins in basal half of wing, outer gradate series apparently absent; FW M forked nearly level with origin of RP2, approximately at level of A2 termination, MA few branched, MP pectinately branched (5–6 branches); FW CuA (3–4 branches) and CuP (11–12 branches) both pectinately branched; A1 elongate, strongly pectinate (≥13 branches) with some irregular end-twigging; A2 strongly pectinate (7 branches); A3 with a few pectinate branches; HW unknown.

Comments. Based on an incomplete forewing fragment, the apparent kempynine affinity of this genus (cf. Osmylinae ) remains tentative. Characters such as the numerous basal crossveins, position of the medial fork and sinuate distal branches of RP suggest that is should be placed in Kempyninae but not conclusively and placement in Osmylinae cannot be ruled out. The wing venation of Kempynosmylus is similar to Arbusella and Jurakempynus , genera similarly lacking the outer gradate series. The status of the genus relative to these genera (and its subfamilial placement) will be clearer once additional, more complete material is discovered.

Figures

FIGURE 11. Kempyninae fossils. A. Jurakempynus sinensis Wang et al. (from Wang et al., 2011a); B. Cretosmylus sibiricus Makarkin (from Makarkin, 1990) (forewing); C. Kempynosmylus zherikhini Makarkin (from Makarkin, 2014) (forewing) (figures not to scale).