Lithosmylidia Riek, 1955

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 12

Lithosmylidia Riek, 1955

( Fig. 5 )

Type species. Lithosmylidia lineata Riek, 1955: 678 (by original designation).

Diagnosis. Forewing medium sized; trichosors present; crossveins poorly preserved in wings of all specimens; costal area with veinlets widely spaced, mostly simple; subcostal area with sc-r crossveins not detected; crossveins in area between RA and RP few or absent (one detected in one specimen of L. parvula ); RP originating close to wing base, with at least 10 branches, very slightly sinuous distally in some species; crossveins of RP field not well preserved in most material, but apparently restricted to a few random ones proximally and two widely separate and irregular gradate series; M forked close to wing base; CuA irregularly pectinate forked, with branches oblique. CuP dichotomously forked with few branches; 1A pectinate, length variable; 2A deeply forked; 3A short and apparently simple; HW unknown.

Comments. A specimen of L. lineata was considered by Riek (1955) as a hind wing, however Lambkin (1988) instead argued that it represented a forewing. Originally, Lithosmylidia was described by Riek (1955) belonging to Kempyninae , but Makarkin et al. (2014) argued that the genus belonged to Archeosmylidae . Characters such as Sc+RA joining the wing margin before the wing apex and the lack of RP crossveins support its placement in Archeosmylidae ; unfortunately, important information about the hind wing cubital and anal vein fields were not preserved in any specimen.

Included species.

L. baronne Lambkin, 1988 ( Australia) (Middle Triassic)

L. lineata Riek, 1955 ( Australia) (Late Triassic)

L. parvula Riek, 1955 ( Australia) (Late Triassic)

Figures

FIGURE 5. Archeosmylidae. Lithosmylidia lineata Riek, forewing (redrawn after Lambkin, 1988) (trichosors and vestiture omitted).