Scotinoecus ruiles, T, Duniesky Rios & Goloboff, Pablo A., 2012

T, Duniesky Rios & Goloboff, Pablo A., 2012, New species of Chilean Hexathelidae (Araneae, Mygalomorphae), Zootaxa 3422, pp. 32-51 : 38-39

Scotinoecus ruiles sp. nov.

Figures 3–4 ; 14; 19–20

Type. Female Holotype (6807 MNHN), Reserva Natural “Los Ruiles ” [35° 37ʹ 0ʺ S 72° 21ʹ 0ʺ W], Cauquenes, Chile, 25 Feb 1992 (P. Goloboff–N. Platnick–M.Ramírez).

Etimology. The specific epithet is a noun in apposition, taken from the type localilty.

Diagnosis. S. ruiles differs from S. cinereopilosus by its larger size, with a longer and proportionally more slender leg I, the ALS larger (almost as big as the PMS) and with 4 spigots. It differs from S. major by its smaller size, the narrower sternum (width: length ratio, 0.87), and metatarsi I with fewer spines. The anterior tarsi have more spines than in either of those species.

Description (Holotype female). Total length 11.52; cephalothorax length 4.20, width 3.36; pars cephalica length 2.16, width 2.64; ALE larger than rest, with both eye rows recurved, with series of 10 thick setae in front of and 7 behind OQ. Dorsal midline of cephalothorax with series of setae that reach fovea, in front of which are 2 longer ones, and 2 or 3 smaller setae inside pit. Chelicerae with 8 large promarginal teeth, and 30/33 denticles towards basal part of furrow; outer margin with abundant reddish hairs. Labium length 0.48, width 1.00, with 86 cuspules and 15 elongate setae along anterior margin. Palpal coxae with 93 cuspules. Sternum length 2.52, width 2.20, with darker spots along margin, and three pairs of sigilla (posterior ones largest). Opisthosoma length 5.28, light brown, with hairs intermixed with short setae; series of light mottles not reaching center form chevron. Spinnerets long; PLS with basal: medial: apical articles of lengths 1.48: 1.00: 1.28; PMS length 0.76; ALS biarticulate, with basal: apical articles of length 0.48: 0.32, apical one conic, with 4 spigots; PMS and ALS almost of same size. Chaetotaxy: Femora: I–IV, series of 5–8 d along midline (apical thickest); I, 1 p a; palp, 1–1–1–1–1–1 d (apical thickest), 1 p a. Patellae: I, 1 p a; II, 1–1 p sup; III, 0–1–1 r, 0–1–1 p sup; IV, 1 r m; palp, 1 p a. Tibiae: I, 0–1–1–0 P, 1–1– 1V POST, 2 V A, all weak and slender; II, 1–1– 1 V POST, 2–1–0 V ANT; III, 1–1 P, 0–1–1–0 R, 0–1– 1 V ANT, 2–2– 2 V; IV, 1–1 R, 0–0–1– 1 V ANT, 1–1– 1 V, 0–0–1– 1 V POST; palp, 1–1– 1 V ANT, 1 V, 1–1– 1 V POST, 1 P. Metatarsi: I, 1–1– 1 V POST, 1 V A, 0–1– 1 V ANT, with 2 apical preening combs (of 2–3 bristles); II, 1–1–1–1– 1 V POST, 1–1–1–0 D, 1 V A, 1 V B, 0–0–1– 1 V ANT, 1 R A, on two sides of long apical spine, 2 preening combs (of 2–3 bristles); III, 1–1–0–1 R, 1–1–1–1 P, 1–1–1–1– 1 V ANT, 1 V A, 1 V B, 1–1–0– 1 V POST, with 3 apical preening combs (of 4–3–2 bristles); IV, 1–1–0–0–1 R, 1–1–0–1–1 P, 0–1–1–1– 1 V ANT, 1 V A, 1 V M POST, 1 V B, with 2 apical preening combs (of 3 bristles). Tarsi: I, 1–1–1–0 R, 1–1–1–0 P; II, 0–1–1–1/1–1–1–1 R, 0–1–1–1/0– 1–0–1 P; III, 0–1–1–1 P, 0–1–1–1 R; IV, 1–1–1–1/0–1–0–1 P, 0–1–1–1 R; palp, 1–0–2–1 R, 1 V A, 1–1–1–1 P, basal lateral spines longer. Trichobothria: Tarsi I–III 7, IV 9; metatarsi I–III 14, IV 21; tibiae I–III with 2 rows of 8, IV with 2 rows of 9.

Variation in other females. Patellae: III, 1 r m, 0–1–1 p sup. Tibiae: I, 1 P M, 2 V A ANT, 1–1– 1 V POST; II, 1 P M, 2 V A ANT, 1–1–0– 1 V POST; III, 1–1–1 R, 1–1–0 P, 0–1– 2 V ANT, 1 V B, 1–1– 1 V POST; IV, 0–1–1–0 R, 0–1– 2 V ANT, 1 V B, 1 V A POST; palp, 1–1– 2 V ANT, 1–1– 2 V POST. Metatarsi: I, 0–1–0– 1 V ANT, 1 V A, 1 V B, 1–1–1– 1 V POST; II, 0–1–0–1 D, 0–1– 1 V ANT, 1–0– 1 V, 1–1–0– 1 V POST, with three apical preening combs (of 2–2–4 bristles); III, 0–1–1– 1 V ANT, 1–0– 1 V, 1–1–0– 1 V POST, 0–1–0–1 D POST, 1 D A ANT, 0–1– 1–0 P, with 3 apical preening combs (of 4–2–4 bristles); IV, 0–1–1– 1 V POST, 1 V B, 0–1–1– 1 V ANT, 0–1–0–1 D A ANT, 0–1–0–1 P, with 2 apical preening combs (of 5–3 bristles). Tarsi: I, 0–1–1–0 P INF, 1 R INF; II, 0–1–1–0 P, 0–1–1–0 R; III, 0–1–1–0 R, 0–1–1–0 P; IV, 0–1–1 R, 0–1–1 P; palp,: 1–1–1– 1 V ANT, 1 V M, 1–1–1– 1 V POST. Trichobothria: tarsi I–III 5, IV 7; metatarsi I–III 9, IV 13; tibiae with 2 rows of 6.

Male. Unknown.

Other specimens examined. Chile. Región VII (del Maule): Talca: Gil de Vilches [35°33'56"S 70°55'51"W], 7 Feb 1992 (P. Goloboff – N. Platnick – M. Ramírez) 2 Ƥ ( MACN). Cauquenes: Reserva Natural “Los Ruiles ” [35°37ʹ0ʺ S 72°21ʹ 0ʺ W], 25 Feb 1992 (P. Goloboff – N. Platnick – M. Ramírez) 1 Ƥ, 2 juv ( MACN).

Distribution. Known from the provinces of Talca and Cauquenes.

Note. The specimens described here come from different localities and present minor differences in their spermathecae, size, and spination. All have the ALS as large as the PMS. In the absence of additional specimens, we prefer to be conservative and consider that all the specimens are conspecific.

Figures

FIGURES 1 – 6. Female Scotinoecus. Sternum and maxillae, and spinnerets. 1 – 2. S. cinereopilosus. 3 – 4. S. ruiles sp. nov. 5 – 6. S. major sp. nov.