Pareuryaptus cambodgiensis Dubault, Lassalle et Roux, 2008

Fedorenko, D. N., 2024, Notes on Trigonotoma and Pareuryaptus (Coleoptera: Carabidae: Pterostichini) from Vietnam, Russian Entomological Journal 33 (1), pp. 43-71 : 64

6. Pareuryaptus cambodgiensis Dubault, Lassalle et Roux, 2008

Dubault et al., 2008: 200, 207 (‘Angkor’); Roux et al., 2016: 76.

DIAGNOSIS. A medium-sized species defined chiefly by sharply carinate inclination of prosternal process. Mandibles laterally rounded; antennal scape barely shorter than antennomeres 2 to 4 combined; pronotal base impunctate between basal foveae, median line obliterate basally. Elytral microsculpture hardly traceable, consisting of moderately transverse meshes. Internal sac of aedeagus ( Fig. 110 ) with basal bulb hooked (all males examined).

REDESCRIPTION. Aedeagus ( Figs 100–105 , 108–110, 113–115 , 138–139 ): slightly asymmetrical in dorsal/ventral view, with left margin angulate at middle, indistinctly sinuate between the angle and apex, this latter curved barely to the left. Internals sac in lateral view at right angle with apical half of aedeagus; with preapical bulb and preapical vesicle large, distal vesicle small and situated on right side of base of dorsal sclerite; distal part with a low yet distinct dorsoapical bulb.

HABITATS AND HABITS. This species appears to be eurybiotic rather than forest-dwelling. Four specimens examined were taken along forest edges at light, and the other four were hand collected along roads or in a cropfield.

COMMENTS. This species is here subdivided into two subspecies. These have hitherto been recognized as separate species, both being rather different in appearance. But their aedeagi, with internal sacs, are very similar and thus argue for the contrary.

Figures

Figs 106–115. Aedeagus, median lobe with everted and inflated internal sac: 106, 111 — Pareuryaptus aethiops; 107, 112 — P. glastenvalum; 108–109, 113–114 — P. cambodgiensis luangphabangensis from Phong Nha – Kebang NP (108, 113) and Kon Ka Kinh NP (109, 114); 110–115 — P. c. cambodgiensis; 106–110 — dorsal aspect; 111–115 — ventral aspect; bp — proximal bulb; bpa — preapcal bulb; ds — dorsal sclerite; vdd — distal dorsal vesicle; vdp — proximal dorsal vesicle. Scale bars 1 mm. Рис. 106–115. Эдеагус, среднЯЯ долЯ с вывернутым и надутым внутренним мешком: 106, 111 — Pareuryaptus aethiops; 107, 112 — P. glastenvalum; 108–109, 113–114 — P. cambodgiensis luangphabangensis иЗ Нац. парков Phong Nha - Kebang (108, 113) и Kon Ka Kinh (109, 114); 110–115 — P. c. cambodgiensis; 106–110 — сверху; 111–115 — сниЗу; bp — проксимальный пуЗырь; bpa — предвершинный пуЗырь; ds — дорЗальный склерит; vdd — дистальный дорЗальный пуЗырёк; vdp — проксимальный дорЗальный пуЗырёк. Масштаб: 1 мм.

Figs 96–105. Aedeagus, median lobe with everted and inflated internal sac: 96–97 — P. aethiops; 98–99 — P. glastenvalum; 100–103 — P. cambodgiensis luangphabangensis from Phong Nha - Kebang NP (100–101) and Kon Ka Kinh NP (102–103); 104–105 — P. c. cambodgiensis; 96, 98, 100, 102, 104 — left aspect; 97, 99, 101, 103, 105 — right aspect; bp — proximal bulb; bpa — preapcal bulb; ds — dorsal sclerite; vdd — distal dorsal vesicle; vdp — proximal dorsal vesicle. Scale bars 1 mm. Рис. 96–105. Эдеагус, среднЯЯ долЯ с вывернутым и надутым внутренним мешком: 96–97 — P. aethiops; 98–99 — P. glastenvalum; 100–103 — P. cambodgiensis luangphabangensis иЗ Нац. парков Phong Nha - Kebang (100–101) и Kon Ka Kinh (102–103); 104–105 — P. c. cambodgiensis; 96, 98, 100, 102, 104 — слева; 97, 99, 101, 103, 105 — справа; bp — проксимальный пуЗырь; bpa — предвершинный пуЗырь; ds — дорЗальный склерит; vdd — дистальный дорЗальный пуЗырёк; vdp — проксимальный дорЗальный пуЗырёк. Масштаб: 1 мм.

Figs 138–147. Right paramere, left aspect: 138 — Pareuryaptus c. cambodgiensis; 139 — P. c. luangphabangensis; 140 — P. aethiops; 141 — P. glastenvalum; 142 — P. parvus; 143 — P. chalceolus; 144–147 — P. morosus from Thailand (144 — La Lu NP; 145 — Tha Plan NP) и Cat Tien NP (146–147). Scale bar 1 mm. Рис. 138–147. ПраваЯ парамера слева: 138 — Pareuryaptus c. cambodgiensis; 139 — P. c. luangphabangensis; 140 — P. aethiops; 141 — P. glastenvalum; 142 — P. parvus; 143 — P. chalceolus; 144–147 — P. morosus иЗ Таиланда (144 — Нац. парк La Lu; 145 — Нац. парк Tha Plan) и Нац. парка Cat Tien (146–147). Масштаб: 1 мм.