Gratiola L.

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1972, Flora Europaea. Volume 3. Diapensiacea to Myoporaceae, Cambridge: Cambridge University Press : 203

1. Gratiola L. 1

Perennial herbs; leaves opposite. Flowers solitary in the leafaxils; pedicel bearing 2 bracteoles just below the calyx. Calyx deeply and unequally 5-lobed. Corolla with wide tube and somewhat 2-lipped limb; upper lip flat, 2-lobed, the lower slightly larger, 3-lobed. Stamens included, the 2 upper lateral fertile, the other 3 reduced to staminodes. Anther-lobes transverse, surrounded by a membranous expansion of the connective. Stigma 2-lobed. Capsule 4-valved (septicidal and loculicidal); seeds numerous. 1 Plant glabrous; pedicels shorter than subtending leaves 1. officinalis

1 Plant glandular-puberulent at least above; pedicels as long as subtending leaves

2 Leaves linear; corolla 12-18 mm; lower staminodes conspicuous 2. linifolia

2 Leaves lanceolate; corolla 8-12 mm; lower staminodes minute 3. neglecta