Cymbalaria Hill

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1972, Flora Europaea. Volume 3. Diapensiacea to Myoporaceae, Cambridge: Cambridge University Press : 236-237

15. Cymbalaria Hill 1

Herbs, usually short-lived perennials but sometimes behaving as annuals; stems procumbent to decumbent. Leaves reniform to suborbicular, petiolate, palmately veined, entire to palmately lobed. Flowers solitary in the leaf-axils. Calyx deeply 5-lobed; segments somewhat unequal. Corolla, stamens and capsule as in Linaria . Seeds usually fairly numerous, variously tuberculate, ridged or alveolate.

Predominantly chasmophytes, confined as natives to the Mediterranean region and the S. Alps.

Literature: G. Cufodontis, Arch. Bot. (Forlì) 12: 54-81, 135- 158, 233-254 (1936); Pflanzenareale 4: 69-71 (1938); Bot. Not. 1947: 135-156 (1947). A. Chevalier, Bull. Soc. Bot. Fr. 83: 638-653 (1937).

1 Stem and leaves glabrous at maturity

2 Corolla not more than 15 mm; middle and upper intemodes usually about equal

3Spur 1-5-3 mm, about as long as calyx; capsule glabrous; seeds remaining separate 1. muralis

3Spur 4-5 mm, much longer than calyx; capsule glandular-puberulent; seeds of each loculus concrescent to a single mass 2. longipes 2 Corolla 15-30 mm; middle internodes usually much longer than upper 4 Spur 6-9 mm, about twice as long as calyx 5. pallida 4 Spur 4-5 mm, about as long as calyx 6. hepaticifolia 1 Stem and leaves with persistent hairs 5 Corolla 15-30 mm; stem puberulent to pubescent 5. pallida 5 Corolla not more than 15 mm; stem ± villous 6 Capsule hairy 7 Most of the leaves 7- to 9-lobed; calyx ± densely villous

7. pilosa 7 Most of the leaves 3- to 5-lobed; calyx glabrous to sparsely hairy 8. microcalyx 6 Capsule glabrous 8 Stem and leaves puberulent; seeds covered with large,

hemispherical tubercles 4. muelleri 8 Stem and leaves pubescent to villous; seeds rugose or alveolate 9 Most of the leaves 5- to 9-lobed 1. muralis 9 Most of the leaves entire or 3-lobed, the larger sometimes

5-lobed 3. aequitriloba