Philonthus phaeopus, Stephens, 1833

Herman, Taphylininae Iochini Aorothiini Thiini Latyprosopini Taphylinini Mblyopinina Nisolinina Yptiomina Hilonthina Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. V. Staphylinine Group (Part 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2001 (265), pp. 2441-3020 : 2441-3020

phaeopus Stephens, 1833 , see: nigritulus Gravenhorst, 1802 .

philo Smetana, 1984: 140 ( Gabrius ; Type locality: Japan: Nikko N. P., Ryuzu).

- DISTRIBUTION: Japan.

philomimus Schillhammer, 2000 c: 125 ( Gabrius ; Type locality: Russia: Primorskiy Kray: Sikhote-Alin , Biol. Stat., 30 km SE Chuguyevka, 44.05N, 134.12E, 650 m).

- DISTRIBUTION: Russia.

philonthoides Schillhammer, 1992: 64 ( Gabrius ; Type locality: China: Gansu, Meijishan ).

- DISTRIBUTION: China.

picipennis Mäklin, 1852: 313 ( Philonthus ; [preoccupied]; Type locality: insula Sitkha).

— Fauvel, 1874: 237 [= 1874a: 463] ( Philonthus ; synonym of quisquiliarius ).

— Ganglbauer, 1895: 460 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 350 ( Philonthus ; catalog).

— Hatch, 1957: 199 ( Philonthus ; subgenus Gabrius ; characters; British Columbia; Washington; Oregon).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1965: 29 ( Gabrius ; characters; notes; Labrador; St. Pierre; Newfoundland).

— Smetana, 1990: 178 ( Gabrius ; characters).

— Smetana, 1995e: 629 ( Gabrius ; characters; notes; all Provinces and Territories of Canada except Prince Edward Island; Alaska; Arizona; California; Colorado; Connecticut; Idaho; Illinois; Maine; Massachusetts; Michigan; Montana; Nebraska; Nevada; New Hampshire; New Mexico; New Mexico; New York; North Dakota; Oregon; Pennsylvania; South Dakota; Utah; Washington; Wisconsin; Wyoming).

— Herman, 2001: 44 ( Gabrius ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Canada, USA.

decipiens Horn, 1884: 217 ( Philonthus ; [preoccupied]; Type locality: California).

— Bernhauer and Schubert, 1914: 334 ( Philonthus ; catalog).

— Fall, 1926: 150 ( Philonthus ; Alaska).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 629 ( Gabrius ; synonym of picipennis ).

lopezi Hatch, 1957: 199 ( Philonthus ; subgenus Gabrius ; Type locality: Washington:

Lopez Is. Also cited from British Columbia, Oregon, and Idaho).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1965: 57 ( Gabrius ; synonym of decipiens ).

— Smetana, 1995e: 629 ( Gabrius ; synonym of picipennis ).

piger Schillhammer, 1997: 32 ( Gabrius ; Type locality: NW-India [Kashmir], Cachmire, Pahalgam Kolakoi, 3400 m).

— Schillhammer, 2000 c: 128 ( Gabrius ; India).

- DISTRIBUTION: India.

piliger Mulsant and Rey, 1876: 553 [= 1877: 409] ( Gabrius ; Type locality: Corte Saint- François, en Corse).

— Bernhauer and Schubert, 1914: 347 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Scheerpeltz, 1933: 1353 ( Philonthus ; synonym of nigritulus ).

— Coiffait, 1951: 110, 114 ( Gabrius ; characters in key; France).

— Smetana, 1957: 63, 68 ( Gabrius ; characters; England; Scandinavia; Italy; Corsica; Caucasus; Turkey; Germany; Czechoslovakia; Austria).

— Smetana, 1958: 247 ( Gabrius ; characters; ecology; Czechoslovakia).

— Smetana, 1960c: 311 ( Gabrius ; characters; Turkey; Sweden; Denmark; Norway; Germany; Holland; Belgium; England; France; Italy; Corsica; Austria; Czechoslovakia; Yugoslovia; Siberia ; Caucasus).

— Palm, 1963b: 106 ( Philonthus ; subgenus Gabrius ; characters; Sweden).

— Horion, 1965: 190 ( Gabrius ; Europe).

— Coiffait, 1974: 82 ( Gabrius ; characters; Europe; Asia Minor; Caucasus; Daghestan; Sweden; Britain; Italy; France; Corsica).

— Pope, 1977: 30 ( Gabrius ; Britain).

— Petrenko, 1980: 82 ( Gabrius ; Caucasus).

— Tóth, 1984: 87 ( Gabrius ; characters; Hungary).

— Lucht, 1987: 106 ( Gabrius ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Sweden).

— Welch, 1993: 229 ( Gabrius ; ovariole number and ovary structure).

— Smetana, 1993b: 49 ( Gabrius ; Slovakia).

— Ciceroni and Zanetti, 1995: 29 ( Gabrius ; Italy).

— Gerend and Braunert, 1997: 198 ( Gabrius ; notes; Luxembourg).

- DISTRIBUTION: Europe, Russia, Caucasus, Turkey.

stipes Sharp, 1910: 129 ( Gabrius ; Type locality: Anglia: Plymouth; Mickleham; Cambridge ).

— Bernhauer and Schubert, 1914: 356 ( Philonthus ; catalog).

— Gridelli, 1920: 152 ( Gabrius ; characters; England; Daghestan; Caucasus).

— Sainte-Claire Deville, 1926: 119 ( Philonthus ; subgenus Gabrius ; notes; Corsica).

— Scheerpeltz, 1933: 1362 ( Philonthus ; catalog).

— Wüsthoff, 1937: 32 ( Philonthus ; illustration of aedeagus).

— Méquignon, 1944: 20 ( Gabrius ; synonym of piliger ).

— Fagel, 1945: 40 ( Gabrius ; characters; Belgium).

— Smetana, 1952b: 119 ( Gabrius ; characters; Czechoslovakia).

— Smetana, 1953: 121 ( Gabrius ; Turkey).

— Smetana, 1953a: 178 ( Gabrius ; Savinian Alps).

— Smetana, 1957: 63 ( Gabrius ; synonym of piliger ).

— Smetana, 1958: 247 ( Gabrius ; synonym of piliger ).

— Smetana, 1960c: 311 ( Gabrius ; synonym of piliger ).

— Lohse, 1964: 189 ( Gabrius ; synonym of piliger ).

— Pope, 1977: 30 ( Gabrius ; synonym of piliger ).

pilipes Bierig, 1940 , see: chiriquiensis Herman, 2001 .

pisciformis Fauvel, 1874: 242 [= 1874a: 468] ( Philonthus ; Type locality: Var, Entraigues).

— Fauvel, 1874: 340 [= 1874b: 46] ( Philonthus ; Spain).

— Mulsant and Rey, 1876: 542 [= 1877: 398] ( Gabrius ; characters; France).

— Eppelsheim, 1887b: 432 ( Philonthus ; synonym of femoralis ).

— Ganglbauer, 1895: 459 ( Philonthus ; subgenus Gabrius ; characters; France; Spain).

— Bernhauer, 1899: 24 ( Philonthus ; notes; Siebenbürgen; Hungary; Bosnia; Austria).

— Bernhauer and Schubert, 1914: 337 ( Philonthus ; synonym of femoralis ).

— Portevin, 1929: 365 ( Philonthus ; characters; France).

— Scheerpeltz, 1933: 1342 ( Philonthus ; catalog).

— Smetana, 1957: 58 ( Gabrius ; synonym of femoralis ).

— Smetana, 1958: 236 ( Gabrius ; synonym of femoralis ).

— Smetana, 1961: 37 ( Gabrius ; synonym of femoralis ).

— Smetana, 1967b: 156 ( Gabrius ; Spain).

— Coiffait, 1974: 67 ( Gabrius ; characters; Spain; France; Pyrenees).

- DISTRIBUTION: France, Spain.

tibialis Mulsant and Rey, 1876: 540 [= 1877: 396] ( Gabrius ; Type locality: environs de Fréjus et dans le Languedoc).

— Duvivier, 1883: 148 ( Philonthus ; catalog; synonym of pisciformis ).

— Ganglbauer, 1895: 459 ( Philonthus ; synonym of pisciformis ).

— Bernhauer and Schubert, 1914: 337 ( Philonthus ; synonym of femoralis ).

— Coiffait, 1951: 105, 114 ( Gabrius ; cited as valid species; characters in key; France).

— Smetana, 1957: 59 ( Gabrius ; cited as valid species; characters; France; Spain).

— Fagel, 1959a: 96 ( Gabrius ; notes; Spain). — Smetana, 1967b: 156 ( Gabrius ; synonym of pisciformis ).

— Coiffait, 1974: 67 ( Gabrius ; synonym of pisciformis ).

gavoyi Bernhauer, 1928b: 12 ( Philonthus ; subgenus Gabrius ; Type locality: Gavoy in Südfrankreich; Granada in Spanien).

— Scheerpeltz, 1933: 1344 ( Philonthus ; catalog).

— Smetana, 1967b: 156 ( Gabrius ; synonym of pisciformis ).

— Smetana, 1973b: 126 ( Gabrius ; notes; synonym of pisciformis ).

— Coiffait, 1974: 67 ( Gabrius ; synonym of pisciformis ).

plumosus Last, 1981: 125 ( Gabrius ; Type locality: Marifuanga, Asaro-Chimbu Divide).

— Last, 1987: 55 ( Gabrius ; Papua New Guinea).

— Schillhammer, 1997: 64 ( Gabrius ; characters; New Guinea).

- DISTRIBUTION: New Guinea.

praedictus Schillhammer, 1997: 57 ( Gabrius ; Type locality: Nepal: 328 Panchthar Distr. , Paniporua, 2300 m).

- DISTRIBUTION: Nepal.

primigenius Joy, 1913: 25 ( Gabrius ; Type locality: N.W. Spain, Pontevedra ).

— Scheerpeltz, 1933: 1367 ( Philonthus ; synonym of velox ).

— Smetana, 1957: 65 ( Gabrius ; synonym of velox ).

— Smetana, 1958: 251 ( Gabrius ; synonym of velox ).

— Smetana, 1960c: 331 ( Gabrius ; characters; Spain; North Africa; France).

— Smetana, 1967b: 159 ( Gabrius ; cited as promigenius; Spain).

— Coiffait, 1974: 60 ( Gabrius ; characters; North Africa; Iberian Peninsula; France).

— Pope, 1977: 30 ( Gabrius ; synonym of velox ).

— Smetana, 1995e: 627 ( Gabrius ; valid species).

— Outerelo, Gamarra, and Salgado, 1998: 129 ( Gabrius ; collected from cave; Spain).

- DISTRIBUTION: France, Spain, Morocco.

mauretanicus Peyerimhoff, 1937a: 266 ( Philonthus ; subgenus Gabrius ; Type locality: Causse du Mahmel, Aurès, vers 1800 m).

— Coiffait, 1951: 112, 115 ( Gabrius ; characters in key; France).

— Smetana, 1957: 65 ( Gabrius ; characters; North Africa; France).

— Smetana, 1960c: 331 ( Gabrius ; synonym of primigenius ).

— Smetana, 1967b: 159 ( Gabrius ; synonym of primigenius ).

— Coiffait, 1973a: 274 ( Gabrius ; Morocco).

— Coiffait, 1974: 60 ( Gabrius ; synonym of primigenius ).

propinquus Cameron, 1933d: 389 ( Philonthus ; subgenus Gabrius ; [preoccupied]; Type locality: Sumatra: Gunung Singgalang, 1000 m; [Note: See lectotype designation by Schillhammer, 1997]).

— Cameron, 1937: 6 ( Philonthus ; Java).

— Schillhammer, 1997: 49 ( Gabrius ; lectotype designation; characters; Sumatra; Java; Bali) .

— Schillhammer, 2000 c: 132 ( Gabrius ; Java; Bali).

— Herman, 2001: 44 ( Gabrius ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Indonesia.

pseudimitator Schillhammer, 1997: 46 ( Gabrius ; Type locality: Thailand: Chiang Mai,

Doi Suthep, 1550 m).

- DISTRIBUTION: Thailand.

pseudopennatus Cameron, 1945c: 716 ( Philonthus ; subgenus Gabrius ; Type locality:

South Africa: Simon’s Town).

— Tottenham, 1956: 305 ( Gabrius ; characters in key).

— Tottenham, 1956b: 207, 208 ( Gabrius ; characters).

- DISTRIBUTION: South Africa.

pubens Mulsant and Rey, 1878 , see: Gabronthus .

turneri Cameron, 1951c: 403 ( Philonthus ; subgenus Gabrius ; [preoccupied]; Type locality: Kenya Colony: Londiani).

— Schillhammer, 1997: 82 ( Gabrius ).

— Herman, 2001: 44 ( Gabrius ; synonym of pullatus ).

punctatellus Horn, 1884: 215 ( Philonthus ; [preoccupied]; Type locality: Ft. McLeod Brit. Amer.; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Hudson’s Bay region; Lake Tahoe ).

— Bernhauer and Schubert, 1914: 352 ( Philonthus ; catalog).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 692 ( Gabrius ; subspecies punctatellus ; lectotype designation: Ft. McLeod Brit. Amer.; characters; notes; Alberta; British Columbia; Saskatchewan; Yukon Territory; Colorado; Idaho; Montana; Oregon; Utah; Washington).

— Herman, 2001: 44 ( Gabrius ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Canada, USA.

ottawensis Casey, 1915: 438 ( Philonthus ; Type locality: Canada: Ottawa ; [Note: See lectotype designation by Smetana, 1995e]).

— Scheerpeltz, 1933: 1354 ( Philonthus ; catalog).

— Moore, 1965: 100 ( Gabrius ; cited as ottawaensis).

— Smetana, 1995e: 693, 695 ( Gabrius ; subspecies of punctatellus ; lectotype designation; characters; notes; Manitoba; Ontario; Quebec; Massachusetts; New Hampshire; New York; Wisconsin).

downiei Hatch, 1957: 203 ( Philonthus ; subgenus Gabrius ; Type locality: Washington: Pullman. Also cited from British Columbia and Idaho).

— Moore, 1965: 100 ( Gabrius ; cited as downei).

— Smetana, 1995e: 692 ( Gabrius ; synonym of punctatellus punctatellus ).

wawawai Hatch, 1957: 203 ( Philonthus ; subgenus Gabrius ; Type locality: Washington; Wawawai. Also cited from Oregon and British Columbia).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 692 ( Gabrius ; synonym of punctatellus punctatellus ).

miltiades Smetana, 1995e: 693 , 695 ( Gabrius ; subspecies of punctatellus ; Type locality: New Mexico: Lincoln N.F., 1 mi. SE Cloudcroft, 8750 feet. Also cited from Arizona, Nevada, Nebraska; labelled as intermediate between miltiades and punctatellus : Colorado and Utah).

pujoli Levasseur, 1967 , see: holisinus Fauvel, 1904 .

pumilus Mannerheim, 1830 , see: nigritulus Gravenhorst, 1802 .

pusillus Heer, 1839 , see: nigritulus Gravenhorst, 1802 .

puthzi Schillhammer, 1997: 35 ( Gabrius ; Type locality: Nepal [ Sindhupalchok District ]:

Prov. Bagmati, Malemchi, 2800 m).

- DISTRIBUTION: Nepal.

pygmaeus Stephens, 1833 , see: nigritulus Gravenhorst, 1802 .

pygmaeus Snellen, 1854 , see: nigritulus Gravenhorst, 1802 .

quadriceps Jarrige, 1978: 285 ( Gabrius ; Type locality: Andringitra Est, Anjavidilava, 2000 m). - DISTRIBUTION: Madagascar.

quadripunctatus Schillhammer, 1992: 64 ( Gabrius ; Type locality: China: Gansu, Meijishan ). - DISTRIBUTION: China.

rabigoides Jarrige, 1957: 110 ( Gabrius ; Type locality: Réunion: forêt de Bélouve).

— Lecoq, 1990: 192 ( Gabrius ; Mascarene Islands).

- DISTRIBUTION: Réunion.

ramifer Tottenham, 1956: 303 ( Gabrius ; Type locality: Ruanda: Shangugu , 1500 m).

- DISTRIBUTION: Rwanda. — Fagel, 1955a: 85 ( Gabrius ; Greece).

— Smetana, 1960c: 351 ( Gabrius ; characters from original description; species uncertain).

— Horion, 1965: 195 ( Gabrius ; Austria; Slovenia; Greece; Holland).

— Bordoni, 1973i: 329 ( Gabrius ; characters).

— Coiffait, 1974: 55 ( Gabrius ; characters; Europe; Asia Minor; Caucasus; Pyrenees).

— Smetana, 1977: 794 ( Gabrius ; Turkey).

— Tóth, 1984: 92 ( Gabrius ; characters; Hungary).

— Schillhammer, 1990: 91 ( Gabrius ; characters; notes).

— Smetana, 1993b: 49 ( Gabrius ; Czech Republic; Slovakia).

— Schillhammer, 1993: 326 ( Gabrius ; Austria).

— Ciceroni and Zanetti, 1995: 29 ( Gabrius ; Italy).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Gabrius ; note; Austria).

— Tronquet, 1998: 11 ( Gabrius ; notes; France).

- DISTRIBUTION: Europe, Caucasus, Turkey.

spurius Smetana, 1954b: 178 ( Gabrius ; [Note: This name was validated by Smetana, 1954b; comparative characters were cited and an illustration of the aedeagus was published; the author stated the species was to be described later]; Type locality: Moravia : Prostějov [ Note : Localities taken from Smetana , 1956b: 175. Also cited from Anatolia, Bohemia, Czechoslovakia, Austria, Hungary and the Caucasus).

— Smetana, 1952b: 122 ( Gabrius ; [Note: Misidentified as subnigritulus according to Smetana, 1958: 250]; characters; Czechoslovakia).

— Smetana, 1957: 67, 69 ( Gabrius ; characters; Czechoslovakia; Austria; Hungary; Transylvania; Bulgaria; Caucasus; Turkey; Germany).

— Smetana, 1958: 250 ( Gabrius ; characters; ecology; Czechoslovakia).

— Smetana, 1960c: 323 ( Gabrius ; characters; Caucasus; Turkey; Czechoslovakia; Austria; Hungary; Transylvania; Bulgaria).

— Smetana, 1961a: 44 ( Gabrius ; Bulgaria).

— Lohse, 1964: 191 ( Gabrius ; characters; central Europe).

— Horion, 1965: 196 ( Gabrius ; Caucasus; Turkey; Bulgaria; Romania; Czech Republic; Slovakia; Austria).

— Smetana, 1967b: 159 ( Gabrius ; Spain).

— Coiffait, 1974: 55 ( Gabrius ; synonym of ravasinii ).

— Smetana, 1977: 794 ( Gabrius ; synonym of ravasinii ).

— Lucht, 1987: 106 ( Gabrius ; checklist; Czechoslovakia; Austria).

— Schillhammer, 1990: 91 ( Gabrius ; notes).

— Smetana, 1993b: 49 ( Gabrius ; synonym of ravasinii ).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Gabrius ; synonym of ravasinii ).

spurius Smetana, 1956b: 171 , 175 ( Gabrius ; [Note: Cited as a new species but the name was validated in Smetana, 1954b]; characters; Moravia. Also cited from Anatolia, Bohemia, Austria, Hungary, Transylvania, Bulgaria, and Caucasus).

nigrituloides Coiffait, 1963: 20 ( Gabrius ; [preoccupied]; Type locality: Forêt de Lente, Drôme).

— Coiffait, 1974: 55 ( Gabrius ; characters; France).

— Schillhammer, 1996a: 81 ( Gabrius ; synonym of ravasinii ).

riedeli Schillhammer, 1997: 70 ( Gabrius ; Type locality: Indonesia: N-Sumatra: Kutacane

Gg. Sinabung, 2000 m).

- DISTRIBUTION: Indonesia.

robustus Smetana, 1953a: 168 ( Gabrius ; Type locality: Macedonia: Kistac ).

— Smetana, 1957: 66 ( Gabrius ; characters; Macedonia).

— Smetana, 1960c: 349 ( Gabrius ; characters; Macedonia).

— Coiffait, 1974: 59 ( Gabrius ; characters; Macedonia).

— Schuh, Schillhammer, and Zettel, 1992: 220 ( Gabrius ; characters; notes; Austria).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Gabrius ; characters; Macedonia; Austria).

roubali Smetana, 1952a: 89 ( Gabrius ; Type locality: Rossia mer., Caucasus occ., Adler).

— Smetana, 1960c: 343 ( Gabrius ; characters; Caucasus).

— Coiffait, 1974: 59 ( Gabrius ; characters; Caucasus).

- DISTRIBUTION: Russia.

roubalioides Schillhammer, 1997: 15 ( Gabrius ; Type locality: O-Afghanistan: Do-Schak, Hindukusch, Khinjantal, 2500 m).

- DISTRIBUTION: Afghanistan.

rougemonti Schillhammer, 1997: 51 ( Gabrius ; Type locality: Bali: Penulisan ).

- DISTRIBUTION: Indonesia.

rubripennis Kiesenwetter, 1844 , see: lividipes Baudi, 1848 .

rufocinctus Schillhammer, 1997: 55 ( Gabrius ; Type locality: Nepal [Sindhupalchok District]: Prov. Bagmati, Malemchi , 2800 m).

— Schillhammer, 2000 c: 133 ( Gabrius ; Nepal).

- DISTRIBUTION: Nepal, India.

dubiosus Schillhammer, 1997: 56 ( Gabrius ; subspecies of rufocinctus ; Type locality: India: W. Bengal, Darjeeling dist., Tigerhill. Also cited from Nepal).

— Schillhammer, 2000 c: 133 ( Gabrius ; subspecies of rufocinctus ; Nepal).

rupicola Cameron, 1943: 131 ( Philonthus ; subgenus Gabrius ; Type locality: Darjeeling: Ghum district; [Note: See lectotype designation by Schillhammer, 1997]).

— Schillhammer, 1997: 29 ( Gabrius ; lectotype designation; characters; India; Nepal) .

- DISTRIBUTION: India, Nepal.

sacerdotalis Joy, 1913: 26 ( Gabrius ; Type locality: Caucasus, Swanetia).

— Gridelli, 1920: 149 ( Gabrius ; characters; Caucasus).

— Scheerpeltz, 1933: 1359 ( Philonthus ; catalog).

— Smetana, 1960c: 339 ( Gabrius ; characters; Caucasus).

— Korge, 1971a: 7 ( Gabrius ; Turkey).

— Coiffait, 1974: 62 ( Gabrius ; characters; Caucasus).

- DISTRIBUTION: Turkey, Georgia.

sagittifer Schillhammer, 1997: 60 ( Gabrius ; Type locality: Nepal: [Sindhupalchok District]: Prov. Bagmati, Malemchi Khola near Malemchi , 2100 m. Also cited from India).

— Schillhammer, 2000 c: 133 ( Gabrius ; Nepal).

- DISTRIBUTION: India, Nepal, China.

acutiphallus Schillhammer, 1997: 61 ( Gabrius ; subspecies of sagittifer ; Type locality: China: Yunnan, 100 km W. Baoshan, Gaoligongshan Nat. Res.).

sallustius Smetana, 1995e: 691 ( Gabrius ; Type locality: Oregon: Fly Cr. Valley , SE Umatilla County. Also cited from Washington).

— Hatch, 1957: 202 ( Philonthus ; subgenus Gabrius ; [Note: Misidentified as distans according to Smetana, 1995e: 689, 691]; characters; British Columbia; Washington).

- DISTRIBUTION: Canada, USA.

satanellus Bordoni, 1973 , see: doderoi Gridelli, 1920 .

schawalleri Coiffait, 1982 , see: furtivus Cameron, 1932 .

schawallerianus Schillhammer, 1997: 56 ( Gabrius ; Type locality: Nepal: 328 Panchthar Distr. , Paniporua, 2300 m).

— Schillhammer, 2000 c: 133 ( Gabrius ; Nepal).

- DISTRIBUTION: Nepal.

schillhammeri Shibata, 1996: 146 ( Gabrius ; replacement name for multipunctatus Shibata ). - DISTRIBUTION: Japan.

multipunctatus Shibata, 1991a: 88 ( Gabrius ; [preoccupied]; Type locality: Japan: Yamanashi Pref.: Akazawa Spa, Masuho ).

— Shibata, 1996: 146 ( Gabrius ; synonym of schillhammeri ).

schmidti Schillhammer, 2000 c: 123 ( Gabrius ; Type locality: Nepal Himalaya : Manaslu-Mts., Dudh Pokhari Lekh zw. Simia Kharka and Malamche Kharka, 33–3500 m).

- DISTRIBUTION: Nepal.

schoedli Schillhammer, 1997: 72 ( Gabrius ; Type locality: Indonesia: S-Sulawesi, E. Malino, Kanre Apia, 1400 m).

- DISTRIBUTION: Indonesia.

schwendingeri Schillhammer, 1997: 46 ( Gabrius ; Type locality: Thailand: Mae Hong Son - Doi Chang, 20 km E Pai).

- DISTRIBUTION: Thailand.

scrutandus Schillhammer, 1997: 35 ( Gabrius ; Type locality: Nepal: [Sankhua Sabha District]: Khandbari Distr., Induwa Khola Valley, 2850 m).

- DISTRIBUTION: Nepal.

seattlensis Hatch, 1957: 201 ( Philonthus ; subgenus Gabrius ; Type locality: Washington: Seattle. Also cited from British Columbia and Oregon).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 669 ( Gabrius ; characters; notes; British Columbia; Oregon; Washington).

- DISTRIBUTION: Canada, USA.

semipunctatus Stephens, 1833: 252 ( Gabrius ; Type locality: Suffolk).

— Stephens, 1839: 401 ( Gabrius ; characters; England).

- DISTRIBUTION: Britain.

senegalensis Cameron, 1949a: 320 ( Gabrius ; Type locality: Guinée: Mt. Nimba , alt. 700–

900 m. Also cited from Ivory Coast).

- DISTRIBUTION: Guinea, Ivory Coast.

septempunctatus Cameron, 1930c: 207 ( Philonthus ; subgenus Gabrius ; Type locality: Kobe: Gomo).

— Scheerpeltz, 1933: 1360 ( Philonthus ; catalog).

— Cameron, 1949b: 468 ( Philonthus ; Formosa).

— Smetana, 1984: 122 ( Gabrius ; species unknown; Japan).

- DISTRIBUTION: Japan, Taiwan.

serpentis Schillhammer, 1997: 60 ( Gabrius ; Type locality: Nepal: [Sindhupalchok District]: Prov. Bagmati, Malemchi , 2800 m).

- DISTRIBUTION: Nepal.

sesengeanus Tottenham: [Note: Nomen nudum (Article 13.1.1); Tottenham, 1962: 143 cited this name in Gabrius , attributed it to himself, cited it from Uganda but without characters and without a reference to an earlier description].

sexualis Smetana, 1954: 42 ( Gabrius ; Type locality: Gallia mer., Alpes Maritimes, In. Vesubie. Also cited from Italy).

— Smetana, 1957: 63 ( Gabrius ; characters; France; Italy).

— Smetana, 1960c: 342 ( Gabrius ; characters; France; Italy).

— Bordoni, 1973i: 327, 331 ( Gabrius ; characters; Italy) .

— Coiffait, 1974: 65 ( Gabrius ; characters; Alps; Italy; North Africa).

— Ciceroni and Zanetti, 1995: 29 ( Gabrius ; probable subspecies of coxalus ; Italy).

- DISTRIBUTION: Italy, France, “North Africa”.

sharpianus Cameron, 1930c: 207 ( Philonthus ; subgenus Gabrius ; Type locality: Settsu; Katsuogi: Nagasaki; Miyanoshita).

— Scheerpeltz, 1933: 1360 ( Philonthus ; catalog).

— Smetana, 1960c: 305 ( Gabrius ; characters; Japan).

— Smetana, 1984: 142 ( Gabrius ; characters; Japan).

- DISTRIBUTION: Japan, China.

conicus Bernhauer, 1939d: 99 ( Philonthus ; subgenus Gabrius ; Type locality: Japan: Tsushima. Nordwestl. China: Chinkiang).