Philonthus putiphar, Saulcy, 1864

Herman, Taphylininae Iochini Aorothiini Thiini Latyprosopini Taphylinini Mblyopinina Nisolinina Yptiomina Hilonthina Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. V. Staphylinine Group (Part 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2001 (265), pp. 2441-3020 : 2441-3020

putiphar Saulcy, 1864 , see: Neobisnius .

putridarius Gistel, 1857: 19 ( Philonthus ; [Note: Nomen dubium]; Type locality: Styriae).

- DISTRIBUTION: Austria.

pygmaeus Kraatz, 1859 , see: Gabronthus .

pyrenaeus Kiesenwetter, 1850: 219 ( Philonthus ; Type locality: Pyrenaei).

— Kiesenwetter, 1851a: 414 ( Philonthus ; characters; Pyrenees).

— Fairmaire and Laboulbène, 1856: 519 ( Philonthus ; characters; France).

— Kraatz, 1857d: 574 ( Philonthus ; characters).

— Fauvel, 1865a: 296 [= 1865b: 46] ( Philonthus ; habitat; France).

— Fauvel, 1874: 236 [= 1874a: 462] ( Philonthus ; synonym of frigidus ).

— Mulsant and Rey, 1876: 405 [= 1877: 261] ( Philonthus ; characters; France).

— Ganglbauer, 1895: 448 ( Philonthus ; synonym of frigidus ).

— Bernhauer and Schubert, 1914: 339 ( Philonthus ; variety of frigidus ).

— Smetana, 1958: 186 ( Philonthus ; synonym of frigidus ).

— Smetana, 1959: 150 ( Philonthus ; aberration of frigidus ; characters).

— Coiffait, 1963: 25 ( Philonthus ; valid species; characters; Pyrenees).

— Coiffait, 1967a: 400 ( Philonthus ; characters; Pyrenees).

— Coiffait, 1974: 254 ( Philonthus ; characters; Pyrenees).

— Outerelo, 1976: 266 ( Philonthus ; Spain).

- DISTRIBUTION: France, Spain.

pyropterus Kraatz, 1859 , see: hepaticus Erichson, 1840 .

quadraticeps Boheman, 1858: 28 ( Philonthus ; Type locality: Buenos Ayres).

— Lynch, 1884: 150 ( Philonthus ; characters; Argentina).

— Bernhauer and Schubert, 1914: 352 ( Philonthus ; catalog).

— Frank, 1982: 43 ( Philonthus ; list of parasites).

- DISTRIBUTION: Argentina.

quadraticollis Sharp, 1885 , see: suspectus Erichson, 1840 .

quadratus Kraatz, 1859 , see: Belonuchus .

quadriceps Cameron, 1950 , see: Gabronthus .

quadricollis Horn, 1884: 194 ( Philonthus ; Type locality: Illinois; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from New York to Missouri and Michigan).

— Fauvel, 1889b: 265 ( Philonthus ; synonym of quisquiliarius ).

— Ganglbauer, 1895: 455 ( Philonthus ; synonym of quisquiliarius ).

— Fauvel, 1903d: 271 ( Philonthus ; synonym of quisquiliarius ).

— Blatchley, 1910: 383 ( Philonthus ; characters; Indiana).

— Bernhauer and Schubert, 1914: 352 ( Philonthus ; catalog).

— Notman, 1919: 135 ( Philonthus ; New York).

— Notman, 1920a: 180 ( Philonthus ; New York).

— Hatch, 1957: 190 ( Philonthus ; characters; British Columbia; Washington).

— Smetana, 1995e: 287 ( Philonthus ; lectotype designation: Illinois; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Nova Scotia; Ontario; Quebec; Saskatchewan; Arkansas; California; Colorado; Connecticut; Delaware; Idaho; Illinois; Mississippi; Missouri; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New York; North Carolina; North Dakota; Ohio; Oregon; Pennsylvania; South Dakota; Tennessee; Utah; Vermont; Virginia; Washington; Wisconsin; Wyoming) .

— Downie and Arnett, 1996: 378 ( Philonthus ; characters; USA).

- DISTRIBUTION: Canada, USA.

fraternus Fall, 1926: 149 ( Philonthus ; Type locality: Dawson , Yukon Territory).

— Scheerpeltz, 1933: 1343 ( Philonthus ; catalog).

— Hochhuth, 1849: 141 ( Philonthus ; characters; Iberia).

— Redtenbacher, 1849: 703 ( Philonthus ; characters; Austria).

— Kiesenwetter, 1851a: 416 ( Philonthus ; France).

— Rosenhauer, 1856: 71 ( Philonthus ; Xerez; Cartama).

— Fairmaire and Laboulbène, 1856: 523 ( Philonthus ; characters; France).

— Kraatz, 1857d: 607 ( Philonthus ; characters; Germany).

— Redtenbacher, 1857: 193 ( Philonthus ; characters; Austria).

— Kraatz, 1858a: 61 ( Philonthus ; Greece).

— Peyron, 1858: 426 ( Philonthus ; Turkey).

— Thomson, 1860: 160 ( Philonthus ; characters; Scandinavia).

— Fauvel, 1865a: 297 [= 1865b: 47] ( Philonthus ; habitat; France).

— Fauvel, 1874: 237 [= 1874a: 463] ( Philonthus ; characters; France; Barbary; Egypt; Turkey; Caucasus; Transcaucasia; Japan; USA).

— Sharp, 1874: 44 ( Philonthus ; cited as quisquiliarus; Japan [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Redtenbacher, 1874: 212 ( Philonthus ; characters; Austria).

— Seidlitz, 1875: 276 ( Philonthus ; characters; Baltic region).

— Mulsant and Rey, 1876: 492 [= 1877: 348] ( Philonthus ; characters; France).

— J. Sahlberg, 1876: 18 ( Philonthus ; Finland).

— Fauvel, 1878: 123 [= 1878a: 43] ( Philonthus ; Algeria).

— Fauvel, 1886: 55 [= 1886a: 47] ( Philonthus ; Morocco; Egypt; Algeria).

— Fowler, 1888: 275 ( Philonthus ; characters; Britain).

— Fauvel, 1889b: 265 ( Philonthus ; New Caledonia [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Eppelsheim, 1892: 334 ( Philonthus ; Uzbekistan).

— Ganglbauer , 1895: 455 ( Philonthus ; characters; Palaearctic and Nearctic regions; Africa ; China; Siam; New Caledonia [ Note : According to Tottenham, 1953: 143 specimens from the latter locality are probably misidentified]) .

— Fauvel, 1902b: 110 ( Philonthus ; Barbary; Egypt).

— Fauvel, 1903d: 271 ( Philonthus ; New Caledonia [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Sainte-Claire Deville, 1906: 97 ( Philonthus ; Corsica).

— Montandon, 1908: 80 ( Philonthus ; Romania).

— Poppius, 1909: 26 ( Philonthus ; subgenus Gefyrobius ; Lena Valley, Siberia ).

— Reitter, 1909: 129 ( Philonthus ; characters; Germany).

— Eichelbaum, 1913: 130 ( Philonthus ; Tanzania [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Bernhauer and Schubert, 1914: 352 ( Philonthus ; catalog).

— Johansen, 1914: 411 ( Philonthus ; characters; Denmark).

— Bernhauer, 1915n: 319 ( Philonthus ; German Southwest Africa [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Bernhauer, 1922c: 232 ( Philonthus ; Formosa [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Rambousek, 1922: 1009 ( Philonthus ; Ethiopia; British East Africa [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Lea, 1925: 220, 221 ( Philonthus ; Australian introduction [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Bernhauer, 1926g: 2 ( Philonthus ; Kamchatka region).

— Bernhauer, 1928: 16 ( Philonthus ; Sumatra [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Portevin, 1929: 362 ( Philonthus ; characters; France).

— Cameron, 1932: 123 ( Philonthus ; characters; Ceylon; India [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Scheerpeltz, 1933: 1357 ( Philonthus ; catalog).

— Cameron, 1933d: 387 ( Philonthus ; Sumatra [Note: According to Tottenham, 1953: 143 this is

— Wüsthoff, 1934: 61 ( Philonthus ; illustration of aedeagus).

— Bernhauer, 1936: 4 ( Philonthus ; Mongolia).

— Gridelli, 1934: 74 ( Philonthus ; Karakoram region).

— Cameron, 1937: 5 ( Philonthus ; Java [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Bernhauer and Chapman, 1939: 71 ( Philonthus ; Kenya [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Hafez, 1939 ( Philonthus ; life history; egg, larval, and pupal characters).

— Méquignon, 1942a: 21 ( Philonthus ; Azores).

— Cameron, 1949b: 468 ( Philonthus ; Japan [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— V. Hansen, 1952: 74 ( Philonthus ; subgenus Philonthus ; characters; Denmark).

— Tottenham, 1953: 143 ( Philonthus ; limited to Europe and parts of Asia and North Africa,

not in North America or the Oriental and Australasian regions).

— Smetana, 1953: 119 ( Philonthus ; Turkey).

— Scheerpeltz, 1954: 114 ( Philonthus , cited as Phylonthus; India [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Scheerpeltz, 1956a: 484 ( Philonthus ; Belgian Congo).

— Smetana, 1958: 215 ( Philonthus ; subgenus Philonthus ; characters; ecology; Czechoslovakia).

— Smetana, 1959: 164 ( Philonthus ; cited as quisquiliarus; characters).

— Scheerpeltz, 1960d: 135 ( Philonthus ; Portugal).

— Scheerpeltz, 1961g: 238 ( Philonthus ; Madagascar [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Scheerpeltz, 1961h: 253 ( Philonthus ; Afghanistan).

— Scheerpeltz, 1963: 13 ( Philonthus ; Afghanistan).

— Scheerpeltz, 1963b: 429 ( Philonthus ; Egypt).

— Raianu, 1963: 82 ( Philonthus ; Romania).

— Palm, 1963b: 91 ( Philonthus ; characters; Sweden; Denmark; Norway; Finland).

— Lohse, 1964: 184 ( Philonthus ; subgenus Philonthus ; characters; central Europe).

— Smetana, 1964d: 76 ( Philonthus ; subgenus Philonthus ; Czechoslovakia).

— Focarile, 1964: 59 ( Philonthus ; Italy).

— Scheerpeltz, 1965: 107 ( Philonthus ; Burma [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Scheerpeltz, 1965b: 402 ( Philonthus ; Sicily).

— Scheerpeltz, 1965c: 506 ( Philonthus ; Sicily).

— Horion, 1965: 166 ( Philonthus ; widespread in central and southern Europe).

— Coiffait, 1967a: 424 ( Philonthus ; characters; Europe).

— Smetana, 1967d: 208 ( Philonthus ; Mongolia).

— Smetana, 1967i: 555 ( Philonthus ; Turkey) .

— Raianu, 1968a: 467 ( Philonthus ; Romania).

— Szujecki, 1968a: 731 ( Philonthus ; Poland).

— Coiffait, 1970d: 143 ( Philonthus ; Uzbekistan).

— Coiffait, 1973a: 274 ( Philonthus ; Morocco).

— Moore and Legner, 1974a: 39 ( Philonthus ; spread by commerce).

— Coiffait, 1974: 199 ( Philonthus ; characters; Europe; Britain; Scandinavia; Asia Minor; Lebanon; North Africa).

— Scheerpeltz, 1974: 151 ( Philonthus ; South Africa [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Scheerpeltz, 1974c: 22 ( Philonthus ; Sudan; Ethiopia [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Bordoni, 1975a: 153 ( Philonthus ; Afghanistan).

— Osella and Zanetti, 1975: 109 ( Philonthus ; collected from nest of Talpa europaea ; notes; Italy).

— Watanabe and Shibata, 1976: 332 ( Philonthus ; Malaysia [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).

— Scheerpeltz, 1976a: 89 ( Philonthus ; Nepal).

— Pace and Zanetti, 1977: 298 ( Philonthus ; Italy).

— Pope, 1977: 29 ( Philonthus ; Britain).

— Hinton, 1981: 661 ( Philonthus ; characters of egg).

— Tronquet, 1981: 71 ( Philonthus ; cited as quisquilarius; Afghanistan).

— Coiffait, 1982a: 232 ( Philonthus ; Nepal).

— Frank, 1982: 43 ( Philonthus ; list of parasites).

— Boller, 1983: 118 ( Philonthus ; larval characters).

— Tóth, 1984: 74 ( Philonthus ; cited as quisquilarius; characters; Hungary).

— Lucht, 1987: 105 ( Philonthus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Nekulisianu, Ostafichuk and Tsygankova, 1988 ( Philonthus ; larval characters; biological notes).

— Borges, 1990: Table II ( Philonthus ; Azores) .

— Aranda and Outerelo, 1991: 121 ( Philonthus ; characters).

— Pilon and Zanetti, 1991: 64 ( Philonthus ; Italy).

— J. Li, 1993: 37 ( Philonthus ; China).

— Smetana, 1993b: 48 ( Philonthus ; subgenus Philonthus ; Czech Republic; Slovakia).

— Hernández, Outerelo, and Gamarra, 1994: 192 ( Philonthus ; Canary Islands).

— Yadav, Chand, and Yadav, 1995: 6 ( Philonthus ; morphology of mitotic and meiotic chromosomes).

— Ciceroni and Zanetti, 1995: 28 ( Philonthus ; Italy).

— Smetana, 1995e: 291 ( Philonthus ; does not occur in North America).

— Mikat, Fremuth, and Prouza, 1997: 111 ( Philonthus ; Czech Republic).

— Owen, 1997a: 149 ( Philonthus ; collecting sites; Ireland).

— Duverger, 1998 ( Philonthus ; host to three species of Laboulbeniaceae ).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Philonthus ; characters).

- DISTRIBUTION: Egypt, Algeria, Morocco, Azores, Europe, Russia, Caucasus, Turkey, Syria, Afghanistan, Uzbekistan, Nepal, Mongolia, China, “Karakorum region”. Note: According to Schillhammer (personal communication) the records for the Oriental region cited above refer to P. variipennis and P. bassus .

inquinatus Stephens, 1832: 223 ( Quedius ; Type locality: Suffolk).

— Curtis, 1837: pl. 638 ( Quedius ; Type locality: Britain).

— Stephens, 1839: 392 ( Quedius ; characters; England).

— G. Waterhouse, 1858: 25 ( Philonthus ; synonym of quisquiliarius ).

— Fauvel, 1874: 237 [= 1874a: 463] ( Philonthus ; variety of quisquiliarius ).

— Ganglbauer, 1895: 455 ( Philonthus ; variety of quisquiliarius ).

— Fauvel, 1903: 159 ( Philonthus ; variety of quisquiliarius ; India).

— Fauvel, 1904a: 59 ( Philonthus ; variety of quisquiliarius ; Belgaum; Persia; Europe; India; Japan; Malaysia; Celebes).

— Bernhauer and Schubert, 1914: 352 ( Philonthus ; variety of punctus ).

— Cameron, 1932: 123 ( Philonthus ; synonym of quisquiliarius ).

— Scheerpeltz, 1933: 1358 ( Philonthus ; aberration of quisquiliarius ).

— Smetana, 1953: 119 ( Philonthus ; variety of quisquiliarius ; Turkey).

— Scheerpeltz, 1954: 114 ( Philonthus , cited as Phylonthus; variety of quisquiliarius ; Ceylon).

— Smetana, 1958: 215 ( Philonthus ; aberration of quisquiliarius ).

— Smetana, 1959: 164 ( Philonthus ; aberration of quisquiliarius ; characters).

— Scheerpeltz, 1961h: 253 ( Philonthus ; aberration of quisquiliarius ; Afghanistan).

— Smetana, 1964d: 76 ( Philonthus ; subgenus Philonthus ; aberration of quisquiliarius ; Czechoslovakia).

— Scheerpeltz, 1965: 107 ( Philonthus ; variety of quisquiliarius ; Burma).

— Coiffait, 1967a: 424 ( Philonthus ; aberration of quisquiliarius ; characters).

— Smetana, 1967i: 555 ( Philonthus ; aberration of quisquiliarius ; Turkey).

— Coiffait, 1974: 199 ( Philonthus ; characters; variety of quisquiliarius ).

— Pope, 1977: 29 ( Philonthus ; synonym of quisquiliarius ).

— Tronquet, 1981: 71 ( Philonthus ; variety of quisquiliarius ; Afghanistan).