Toktokkus, Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith, 2021

Kamiński, Marcin J., Gearner, Olivia M., Kanda, Kojun, Swichtenberg, Kali, Purchart, Luboš & Smith, Aaron D., 2021, First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group, Zoological Journal of the Linnean Society 191, pp. 883-901 : 894

KEY TO THE SPECIES OF THE GENUS TOKTOKKUS

1. Apex of elytra not sloped (dorsal view) and depressed with clear margin ( Fig. 3B)............... T. congolensis

- Apex of elytra sloped (dorsal view) and flat to slightly convex (e.g. Fig. 3A) ...............................................2

2. Margin of prosternal collar expanded and folded out into a large lip ( Fig. 3C); elytra round, almost heart- shaped ( Fig. 3C); declivous portion of elytra with little to no tubercles..................................... T. tschinkeli

- Margin of prosternal collar not expanded, only occasionally folded out; elytra round or elongate; declivous portion of elytra with tuberculate rows .........................................................................................................3

3. Disc of pronotum with prominent punctures ( Fig. 3D); gold setae present on elytra ( Fig. 3D)..... T. herero

- Disc of pronotum only with micropunctures; elytra not covered with setae................................................4

4. Tubercles on elytral sides round/globular, vertically direct ( Fig. 2B, C) ......................................................5

- Tubercles on elytral sides pointed, directed posteriorly (e.g. Fig. 3F) ..........................................................6

5. Elytral tubercles dense (1.0–1.5 diameters apart), laterally reaching humerus; elytra round ( Fig. 2B) ........................................................................................................................................................... T. mulleri

- Elytral tubercles sparse (2–3 diameters apart), laterally terminating prior to humerus; elytra elongate to round ( Fig. 2C)................................................................................................................................ T. schultzei

6. Elytral tubercles sparse (4–6 lateral rows), deeply angled...........................................................................7

- Elytral tubercles dense (more than 6 lateral rows), slightly angled ( Fig. 2B–F).........................................8

7. Elytral tubercles distinct, not confluent, nearly reaching humeri ( Fig. 3A) ................................ T. barclayi

- Elytral tubercles small and short, confluent into rows, terminating well before humeri ( Fig. 2A) ...................................................................................................................................................... T. mashunus

8. Microtubercles present between tuberculate rows (e.g. Fig. 3E), tuberculate rows rarely elevated on ridges ...............................................................................................................................................................9

- Elytral tubercles all relatively the same size, no microtubercles present between rows, tuberculate rows often elevated on ridges ................................................................................................................................10

9. Body size fairly small (18.0–23.0 mm); elytral tubercles relatively dense, almost confluent; tubercles relatively short .................................................................................................................... T. tuberculipennis

- Body size medium to large (28.0–32.0 mm); elytral tubercles less dense, only occasionally confluent; tubercles taller ................................................................................................................................ T. makuya

10. Elytral tubercles large, often confluent ( Fig. 3F); tuberculate rows extend over humeri to scutellum; humeri prominent; disk of elytra where smooth often flat ( Fig. 3F) ......................................... T. waclawae

- Tubercles medium sized ( Fig. 2E, F), not confluent; tuberculate rows end at humeri; humeri not prominent; disc of elytra where smooth generally convex .................................................................................... T. vialis