Rhagio vermileo, (Linnaeus, 1758)

Bueno, Gabriel M., Kehlmaier, Christian & Santos, Charles Morphy D., 2021, A worldwide catalog of the Vermileonidae (Diptera: Brachycera), Zootaxa 5060 (4), pp. 489-514 : 502

vermileo ( Linnaeus, 1758)

Musca vermileo Linnaeus, 1758

Rhagio vermileo Fabricius, 1775

Nemotelus vermileo De Geer, 1776

Leptis vermileo Fabricius, 1805

Vermileo degeerii Macquart, 1834

Psammorycter vermileo Blanchard, 1840

Apogon dufouri Perris, 1852

Holotype: sex not indicated in orig. des., “southern France ” (unknown deposit).

Dist.: occurring from eastern mainland Spain to Croatia, including France, Italy, Switzerland, Germany, Austria and Slovenia ( Beuk & Pape 2013; Kehlmaier 2021). SPAIN: Cabrils {41°31’N 2°22’E}; FRANCE: type was assigned to occur in “southern France”, other cited locations are Bouches-du-Rhône, La Ciotat {43°10’N 5°36’E}, Nantes [47°14’19.62’’N 1°33’17.60’’E], Les Eyzeis-de-Tayac-Sireuil {44°56’N 1°02’E}, Les Grandes Landes {45°29’N 1°01’E} and Charnay {45°53’N 4°39’E}. ITALY: the species occurs in the northern half of the mainland and Sardinia ( Kehlmaier 2021); there is also a report from Giglia Island {42°21’N 10°53’E}; historic records from southern Italy including Sardinia and Sicily need to be re-evaluated; AUSTRIA: reported from Linz {48°17’N 14°18’E}; CROATIA: reported from Rovinj {45°04’N 13°40’E}, Premantura {44°47’N 13°54’E}, Lošinj {44°40’N 14°22’E}, Podaca {43°08’N 17°19’E} Dubrovnik-Neretva, Mljet {42°44’N 17°32’E} and Baška Voda {44°58’N 14°47’E}; SLOVENIA: from Osp {45°34’N 13°51’E}, Piran {45°31’N 13°34’E} and Fiesta {45°31’N 13°34’E}; SWITZERLAND: from Montorge (Valais) {46°13’N 7°20’E}; GERMANY: Weil am Rhein, gravel pit Käppelin [47°35’08.3’’N 7°37’52.6’’E]. Historic records from Bulgaria and Greece need to be re-evaluated.

Refs.: De Geer, 1752: 178–191, 261–265 (larv. biol.), Tab.V (figs.); Réaumur, 1753 (larv. biol.); Linnaeus, 1758: 590 (des.); Fabricius, 1775: 762 (redes.) as Rhagio vermile o; De Geer, 1776 (larv. biol.) as Nemotelus vermile o; Fabricius, 1781: 419–420 (as Rhagio vermileo ); Fabricius, 1787: 332 (as Rhagio vermileo ); Fabricius, 1805: 70 (cat.) as Leptis vermile o; Lamarck, 1816: 423 (cat.) as Rhagio vermile o; Cuvier, 1817: 618–619 (as Rhagio vermileo ); Meigen, 1820: 96–97 (as Leptis vermileo ); De Romand, 1833: 498–499 (larv. biol. as Leptis vermileo ); Macquart 1834: 428–429 (redes.) as V. degeeri i; Blanchart, 1840: 603 (redes.), as Psammorycter vermile o; Perris, 1852: 199 (redes.) as Apogon dufour i; Rondani, 1856: 153 (cat.); Von Siebold, 1861 (larv.) biol. as Leptis vermile o; Schiner, 1862: 170–171 (as Apogon dufourii ); Courtiller, 1867: 71, 73–75 (larv. biol.), 72 (figs.) as Leptis vermileo ; Palm, 1869: 417 (dist.) as V. degeer i; Brauer, 1883a: 43–44 (as Vermileo degeeri ); Brauer, 1883b: 114 (as Vermileo degeeri ); Williston, 1886: 107 (note about spp. nomenclature); Mik, 1887: 331 (syn. discussion); Röder, 1892 (syn. discussion); Coucke, 1893: 130 (as Vermileo degeeri ); Mik, 1894 (syn. discussion); Griffini, 1895: 1–2 (dist.) as Psammorcyte r; Bezzi, 1898: 28 (dist. as Psammorcyter vermileo ); Kertész, 1908: 297–298 (cat.); Verral, 1909: 257 (figs.) as V. degeer i; Bequaert, 1922: 277–278 (larv. biol.); Lindner, 1925: 6 (cat., fig.); Séguy, 1926: 91, 93 (figs.), 92 (des., larv. des.); Wheeler, 1930: 144–146 (nomenclature com.), 146–149 (dist.), 150–155 (larv. biol.); Edwards, 1935: 56 (fig.); Buchner, 1940 (biol., larv., biol.); Brauns, 1954 (pupae des.); Le Faucheux, 1961 (biol.); Tsacas, 1962: 200–213 (larv. des.), 201–203, 205–207, 209–212, 214, 225, pls. II, XIII–XVI (figs.), 213–215 (larv. biol.), 216–226 (comparisons to Chrysopilus and Rhagio larv.); Hemmingsen & Nielsen, 1971: 178–181 (biol.); Chassagnard & Tsacas, 1974: 14–26 (larv. morphology, comparison to Chrysopilus auratus ), 15, 18, 20, 23–24, 26 (figs.); Majer, 1988: 34 (cat.); Ebejer, 1995: 87 (wrong species report, probably V. immaculatus ); Thompson & Pont, 1993: 132 (list); Ludwig et al., 1996: 10–11 (larv. des.), 11–12 (larv. biol.); Nagatomi et al., 1999: 20 (key), 23 (cat.); Ovtshinnikova, 2000: 437–438 (male gen.); Carles-Tolrá 2001: 59 (dist.); Ludwig et al., 2001: 92 (larv. des.); Carles-Tolrá & Baéz, 2002: 118 (cat.); Marchesi et al., 2005: 68 (dist.); Devetak, 2008a: 285 (dist., larv. biol.); Devetak, 2008b: 632–634 (larv. biol.); Kerr, 2010: 49,51 (figs.), 58–60, 64 (phyl.); Doczkal, 2017: 826 (dist.); Woodley & Swart, 2017: 866, 868 (figs.); Papp & Soltész, 2019: 5–7 (larv. biol.), 6 (dist.); Hubenov, 2019 (dist.), Kehlmaier, 2021: 342 (phyl., dist.).