Tobantilla aleatrix, Williams, Kevin A., Brothers, Denis J. & Pitts, James P., 2011

Williams, Kevin A., Brothers, Denis J. & Pitts, James P., 2011, New species of Tobantilla Casal, 1965 and a new genus and species, Gogoltilla chichikovi gen. et sp. nov., from Argentina (Hymenoptera: Mutillidae), Zootaxa 3064, pp. 41-68 : 44-45

Key to species of Tobantilla

Males (unknown for T. aleatrix sp. nov., T. charrasca Casal, T. frigidula (Cresson) , T. kolasma sp. nov., T. krima sp. nov., T. montonera Casal )

1. Anteromedial margin of clypeus flat, shallowly emarginate ( Fig. 36 )............................................. 2

- Anteromedial margin of clypeus with single raised tubercle ( Fig. 35 )............................................. 3

2. Base of T2 with patch of appressed white setae ( Fig. 16 )......................................... T. drosos sp. nov.

- T2 having asetose interspaces and sparse erect white setae throughout disc ( Fig. 17 )............... T. ephemeros sp. nov.

3. Metasoma entirely black except for second segment ( Fig. 15 ); legs blackish, concolorous with pleura... T. andrikos sp. nov.

- Metasoma entirely yellowish-brown except segments six and seven infuscated ( Fig. 18 ); legs yellowish-brown, much lighter than dark brown to black pleura............................................................ T. xouthos sp. nov.

Females (unknown for T. andrikos sp. nov., T. drosos sp. nov., T. ephemeros sp. nov., T. xouthos sp. nov.)

1. Head and mesosoma dark red to black ( Figs 3 , 9 ); pygidium having single longitudinal medial carina apical to small pygidial plate ( Figs 13 , 37 )..................................................................... T. frigidula (Cresson)

- Head and mesosoma pale brown to orange (e.g. Figs 1, 2 ); large pygidial plate well defined by lateral carinae, reaching apex of pygidium (e.g. Figs 14 , 38 )............................................................................. 2

2. Scutellar scale connected with carinae separating dorsal and posterior propodeal faces, forming broad continuous arcuate ridge ( Fig. 4 ); posterior face of propodeum clothed with dense black setae ( Fig. 10 )....................... T. kolasma sp. nov.

- Scutellar scale relatively narrow, distinct from carinae separating dorsal and posterior propodeal faces (e.g. Figs 2, 6 ); posterior face of propodeum clothed with pale silver, golden, or brown setae (e.g. Figs 8, 12 )................................. 3

3. T2 having only one pair of yellow integumental spots ( Fig. 5 ); pronotum clothed with black setae, contrasting with silver mesonotal setae ( Fig. 5 )................................................................... T. krima sp. nov.

- T2 having two pairs of yellow integumental spots, sometimes with anterior and posterior spots confluent ( Figs 1, 2, 6 ); mesosomal dorsum clothed with concolorous pale golden setae, sometimes with scattered brown setae on posterior portion of mesonotum ( Figs 1, 2, 6 ).............................................................................. 4

4. T4 clothed entirely with silver setae; head about 1.2 × width of pronotum; T2 with anterior and posterior spots confluent or scarcely separated ( Fig. 6 )............................................................... T. montonera Casal

- T4 having black setae, at least basally; head about 1.1 × width of pronotum; T2 with anterior and posterior spots distinctly separated ( Figs 1, 2 )...................................................................................... 5

5. Mesosomal width slightly greater than length, mesothorax as wide as pronotum, propodeum parallel-sided and flattened posteriorly with posterolateral angle abrupt ( Fig. 2 ); femora darkened medially; flagellomere two only slightly longer than pedicel..................................................................................... T. charrasca Casal

- Mesosomal length slightly greater than width, mesothorax narrower than pronotum, propodeum narrowing posterad and weakly convex posteriorly with posterolateral angle rounded ( Fig. 1 ); femora entirely light orange brown; flagellomere two about one-third longer than pedicel......................................................... T. aleatrix sp. nov.

Figures

FIGURES 19 – 38. Tobantilla species. 19 – 34. Male genitalia: dorsal view (left), ventral view (middle), lateral view of penis valve (above), and lateral view with penis valve removed (right). 19 – 22. T. andrikos sp. nov.; 23 – 26. T. drosos sp. nov.; 27 – 30. T. ephemeros sp. nov.; 31 – 34. T. xouthos sp. nov. 35 – 36. Male clypeus. 28. T. andrikos sp. nov.; 29. T. ephemeros sp. nov. 37, 38. Female pygidium (T 6). 37. T. frigidula (Cresson); 38. T. kolasma sp. nov.

FIGURES 15 – 18. Tobantilla species, males, habitus, lateral view. 15. T. andrikos sp. nov., holotype; 16. T. drosos sp. nov., holotype; 17. T. ephemeros sp. nov.; 18. T. xouthos sp. nov., holotype.

FIGURES 1 – 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal.

FIGURES 7 – 14. Tobantilla species, females. 7 - 12. Habitus, lateral view. 7. T. aleatrix sp. nov., holotype; 8. T. charrasca Casal; 9. T. frigidula (Cresson); 10. T. kolasma sp. nov., holotype; 11. T. krima sp. nov., holotype; 12. T. montonera Casal. 13, 14. Pygidium (T 6). 13. T. frigidula; 14. T. aleatrix.