Omalium caesum Gravenhorst, 1806

Shavrin, Alexey V. & Khachikov, Eduard A., 2023, Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae, Omaliini), Zootaxa 5227 (4), pp. 443-458 : 444-447

Omalium caesum Gravenhorst, 1806

( Figs. 1 , 11–12 , 33 )

Omalium caesum Gravenhorst, 1806: 209

Material examined. RUSSIA: ROSTOV AReA : 1 ♀: Sholokhovskiy District, Elanskaya St. 24– 26.06.1999. E.A. Khachikov leg. (cKh) ; 1 ♀: same data, Veshenskaya. 05.09.2002 (cKh) ; 1 ♀: same data. 26.06.2004 (cKh); VOLGOGRAD AReA: 1 ♁: Elton Lake. 01– 10.08.2006. K.V. Makarov & A.V. Matalin leg. ( MSPU); KRASNODAR TeRRITORY: 1 ♀: Kustshevskiy District, Pervomaiskiy. Forest belt, sieve+sifter. 04.06.2009. L.B. Rybalov leg. ( ZMM); 1 ♀: Ubinskaya. 25.05.1984. I.A. Ushakov leg. ( ZMM); 1 ♀: Temnolesskaya. 25.06.1999. V.V. Grebennikov leg. (cSh); STAVROPOL TeRRITORY: 1 ♀: Pyatigorsk, Mashuk Mt. 17.06.1993. E.A. Khachikov leg. (cKh); 1 ♀: Essentuki. 10.03.2000. E.A. Khachikov leg. (cKh); 1 ♀: Zheleznovodsk, Razvalka. 08.06.1985. A.B. Ryvkin leg. (cR); ADYGeA: 2 ♁♁: Maykop. 06.10.2009. A.R. Bibin leg. (cKh); 1 ♁, 1 ♀: Maykop, Khanskaya, right bank of Belaya River, 44°41′03.54′′N 39°55′41.25′′E. Flood-plain forest. 15.05.2014. E.A. Khachikov leg. (cKh); 1 ♁: same data, 04.06– 02.07.2009 (cKh); 4 ♁♁, 6 ♀♀: Khanskaya. 19.06.2015. E.A. Khachikov leg. (cSh); 2 ♁♁, 3 ♀♀: same data. 16.05.2015 (cKh, cSh); 1 ♀: Mezmay. 19– 25.06.1996. E.A. Khachikov leg. (cKh) 1 ♁: Caucasus Nature Reserve, Guzeripl. 21.06.1986. I.A. Ushakov leg. ( ZMM); 2 ♁♁, 2 ♀♀: same data, 18– 23.05.2015. E.A. Khachikov leg. (cSh); 3 ♁♁, 1 ♀: same data, 22– 23.08.2015 (cKh, cSh); 5 ♁♁, 2 ♀♀: 05.10.2015 (cKh, cSh); 2 ♁♁, 5 ♀♀: same data, Abago (cKh, cSh); 1 ♁, 1 ♀: Abago. 1000 m a.s.l. Forest with Fagus , Abies , motley grass. 02– 20.05.2009. S. Chumachenko leg. (cKh); 3 ♁♁: Abago, Klenovaya Roshcha. 1750 m a.s.l. 29.08– 28.09.2009. S. Chumachenko leg. (cKh); 1 ♀: 11 km Guzeripl-Abago road. 27.07.2014. E.A. Khachikov leg. (cKh); 4 ♁♁, 2 ♀♀: same data. 2015 (cSh); KARACHAY-CHeRKeSSIA: 1 ♁, 1 ♀: Teberda, Dombay. 07.2010. E.A. Khachikov leg. (cSh); 1 ♀: Teberda State Nature Biosphere Reserve,Alibek Gorge, Dvuyazychniy Glacier. 12.07.1996. E.A. Khachikov leg. (cKh); 1 ♀: same Nature Reserve, Malaya Khatipara Mt. 1900 m a.s.l. 29.05.1985. A.B. Ryvkin leg. (cR); KABARDINO-BALKARIA: 1 ♁, 1 ♀: Kabardino-Balkaria, near Elbrus Mt. 2100–2350 m a.s.l., Betula and Pinus forest. 15– 19.08.1989. S.I. Golovatch leg. (cSh); NORTH OSSeTIA-ALANIA: 1 ♀: bank of Terek River near Elkhotovo. 29.07.1984. S.K. Alexeev leg. ( ZMM); 63 ♁♁, 43 ♀♀: Ardon River, Zincar vill. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); 6 ♁♁, 6 ♀♀: Ardon River, Zincar. Taxus , traps. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); 16 ♁♁, 4 ♀♀: Biragzang, southern slope. Traps. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); GEORGIA: SAMeGReLO-ZeMO SVANeTI: 1 ♀: 30 km NE Khaishi, near Kherkhvashi. 1700 m a.s.l., litter near stream. 29.08.1986. A.B. Ryvkin leg. (cR); IMeReTI: 1 ♀: Mukhura, ca. 15 km E of Tkibuli, 700–800 m a.s.l., forest with Castanea , Fagus , Carpinus etc., under bark and stones. 07– 09.05.1987. S.I. Golovatch & K.Yu. Eskov leg. ( ZMM); MTSKHeTA-MTIANeTI: 2 ♁♁: Saguramo Reserve, Zedazeni. 1100–1200 m a.s.l., forest with Fagus , Carpinus , Acer etc,, litter and under bark. 20.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); TBILISI: 9 ♁♁, 12 ♀♀: Tskhneti, 1150–1300 m a.s.l., Fagus , Carpinus , Acer etc. forest, litter. 16, 18.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cR, cSh); KVeMO-KARTLI: 2 ♀♀: Algeti National Park, W of Manglisi. 1400–1500 m a.s.l., forest with Fagus , Picea , Abies etc., in litter and under bark. 16– 18.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); KAKHeTI: 7 ♁♁, 3 ♀♀: Mariamjvari Nature Reserve, NNE of Sagarejo. 1150–1250 m a.s.l., forest with Fagus , Carpinus , Acer , Pinus etc. 13– 14.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM); 1 ♁: N Kvareli. 700–750 m a.s.l., forest with Fagus , Carpinus , Quercus etc., in litter and under bark. 04.05.1987. S.I. Golovatch & K.Yu. Eskov leg. ( ZMM); 1 ♀: Lagodekhi Nature Reserve. 500 m a.s.l. wet litter in forest near river. 05– 06.05.1983. S.I. Golovatch & A.B. Ryvkin leg. ( ZMM); ARMENIA: LORI: 1 ♁, 1 ♀: Odzun W of Alaverdi. 1500–1550 m a.s.l., forest with Quercus , Fagus , Carpinus , etc., litter and under stones. 23– 24.07.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); TAVUSH: 1 ♀: Berdavan, ca 10 km N of Noyemberyan. 900–950 m a.s.l. Forest with Quercus , Carpinus , Acer etc., under stones. 24– 25.05.1987. S.I. Golovatch & K.Yu. Eskov leg. ( ZMM); 2 ♁♁, 1 ♀: Dilijan National Park, Akhnabat valley near Salakh. 1400–1500 m a.s.l., forest with Fagus , Taxus , Carpinus etc., in litter, under bark and stones. 17.04.1983. S.I. Golovatch leg. (cSh, ZMM); 3 ♁♁, 3 ♀♀: W Shamshadyn halfway between Ijewan and Bered. 1500–1600 m a.s.l., forest with Fagus , Carpinus , Acer etc. 26– 27.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM); AZERBAIJAN: SHAKI-ZAQATALA: 4 ♁♁, 2 ♀♀: N Vartashen (Oðuz). 900 m a.s.l, forest with Fagus and Quercus , in litter. 02.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM); 1 ♁: NW above Baş Layisqi, ca. 20 km NNW of Sheki. 1250 m a.s.l., forest with Fagus , Carpinus , Acer etc., litter. 03.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); TURKEY: BURSA: 1 ♁: Uludað. 1900 m a.s.l. 22.07.1969. C. Besuchet leg. ( MHNG); 1 ♁: Uludað, 17 km Bursa. 1100 m a.s.l. 13.05.1976. C. Besuchet & I. Löbl leg. ( MHNG); ZONGULDAK: 1 ♁, 1 ♀: 17 km N Safranbolu. 1000 m a.s.l. 16.05.1976. C. Besuchet & I. Löbl leg. ( MHNG); BOLU: 1 ♀: Abant. 1450 m a.s.l. 25.05.1967. C. Besuchet leg. ( MHNG); MANISA: 2 ♀♀: Salihli. 500 m a.s.l. 29.04.1975. C. Besuchet & I. Löbl leg. ( MHNG); GȔMȔşHANe: 2 ♀♀: Erzincan-Keklit. 2100 m a.sl. 04.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG); ERZINCAN: 1 ♀: Tercan, Euphrate. 1400 m a.s.l. 06.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG); ARTVIN: 2 ♁♁: SW Artvin. 1900 m a.s.l. 09.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG); 2 ♀♀: S Pirnalli, Karkal Daði, 1250 m a.s.l. 11.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG); HATAY: 57 ♁♁, 28 ♀♀: Antakya, Kişlak-Şenköy. 800–850 m a.s.l. 02.05.1978. C. Besuchet & I. Löbl leg. ( MHNG).

Published records. AZERBAIJAN: Leder 1886: 120; GEORGIA: Hochhuth 1849: 204, Zanetti 2002: 56; NAGORNO-KARABAKH: Assing & Schülke 2019: 99; RUSSIA: ADYGeA: Zanetti 2002: 57; DAGeSTAN: Khachikov et al. 2021: 250; KABARDINO-BALKARIA: Khachikov 1998: 6; KRASNODAR TeRRITORY: Roubal 1911: 1; Zanetti 2002: 57; TURKEY: J. Sahlberg 1913: 58; Bordoni 2010: 47.

Distribution. Omalium caesum is widely distributed in the Palaearctic Region: Europe (including British islands), north Africa, Middle Asia, Western and Eastern Siberia ( Schülke & Smetana 2015).

Remarks. Habitus as in Fig. 1 . Aedeagus as in Figs. 11–12 . Female genital segments as in Fig. 33 .

It is here recorded from several regions of SW Russia (Stavropol Territory, Karachay-Cherkessia, North Ossetia-Alania) and Armenia for the first time .

Figures

FIGURES 1–10. Habitus of Omalium: 1—O. caesum (Turkey, Bursa: Uludağ), 2—O. cinnamomeum (Turkey, Antakya: Kişlak- Şenköy), 3—O. excavatum (Russia, Karachay-Cherkessia: Arkhyz), 4—O. falsum (Georgia, Tskhneti), 5—O. littorale (Russia, Adygea: Abago), 6—O. riparium impar (Russia, Krasnodar Territory: Khosta), 7—O. rivulare (Armenia, Alaverdi), 8—O. rugatum (Iran, Zagros Mts.), 9—O. turcicum (Russia, Adygea: Lagonaki), 10—O. wunderlei (Turkey, Gümüşhane: Erzincan- Kelkit). Scale bars: 1.0 mm.

FIGURES 11–18. Aedeagus of Omalium caesum (Figs. 11–12: Turkey, Antakya: Kişlak-Şenköy), O. cinnamomeum (Figs. 13–14: Turkey, Antakya: Kişlak-Şenköy), O. cribriceps (Figs. 15–16: Iran, Lorestan), O. excavatum (Figs. 17–18: Russia, Karachay-Cherkessia: Arkhyz): 11, 13, 15, 17—parameral view, 12, 14, 16, 18— lateral view. Scale bars: 0.1 mm.

FIGURES 33–41. Female genital segment (ventral view) and spermatheca (shown by arrow in Fig. 40) of Omalium: 33—O. caesum (Russia, North Ossetia-Alania: Zincar), 34—O. cinnamomeum (Turkey, Antakya: Kişlak-Şenköy), 35—O. excavatum (Russia, Krasnodar Territory: Caucasus Nature Reserve), 36—O. falsum (Turkey, Rize: Ardeşen), 37—O. littorale (Russia, Volgograd Area: Elton Lake), 38—O. riparium impar (Georgia, Tskhneti), 39—O. rivulare (Russia, Krasnodar Territory: Derbiy), 40—O. rugatum (Russia, North Ossetia-Alania: Zincar), 41—O. turcicum (Russia, Krasnodar Territory, Caucasus Nature Reserve). Scale bars: 0.1 mm.