Fluminicola umbilicatus, Hershler & Liu & Frest & Johannes, 2007

Hershler, Robert, Liu, Hsiu-Ping, Frest, Terrence J. & Johannes, Edward J., 2007, Extensive diversification of pebblesnails (Lithoglyphidae: Fluminicola) in the upper Sacramento River basin, northwestern USA, Zoological Journal of the Linnean Society 149 (3), pp. 371-422 : 407

FLUMINICOLA UMBILICATUS SP. NOV. (HAT CREEK

PEBBLESNAIL)

Type material: Holotype ( Fig. 19A ), USNM 1020705 , Big Spring, tributary of Hat Creek , south of Old Station ( PO) at the south end of Hat Creek Hill , flowing from beneath CA 89 /44 on the south side, Lassen National Forest , Shasta County, California (629540 E, 4500140 N, 1403 m), 24 October 1992 TF, EJ . Paratypes (from same lot), USNM 1020706 .

Referred material: CALIFORNIA. Shasta County: USNM 1020707 , topotypes, 28 August 2001 TF, EJ. – USNM 1020708 , Lost Creek (lower site) in lava field on north side of Wilcox Road , west of Hat Creek Rim , Lassen National Forest (632120 E, 4513600 N, 1141 m), 17 August 1991 TF, EJ, JJ. – USNM 1020709 , ibid., 24 October 1992 TF, EJ. – USNM 1020710 , ibid., 29 August 2001. TF, EJ. – USNM 1020711 , Lost Creek (upper site) in lava field on north side of Wilcox Road , west of Hat Creek Rim , Lassen National Forest (632560 E, 4513140 N, 1141 m), 29 August 2001 TF, EJ. – USNM 1020799 , Lost Creek (upper most site) in lava field on north side of Wilcox Road , west of Hat Creek Rim , inside and west of Lassen National Forest boundary (632730 E, 4513100 N, 1141 m), 3 November 2002 TF, EJ. – USNM 1020716 , Hat Creek on upstream (east) side of CA 89 bridge, both sides of creek at Hat Creek Resort , east of Hat Creek Hill , Old Station Post

Figures

Figure 19. Shells, opercula, and radula of Fluminicola umbilicatus. A, holotype (USNM 1020705). B, USNM 1020706. C, USNM 1020709. D, USNM 1020710. E, USNM 1020711. F, USNM 1020716. G, USNM 1020706. Scales = 1.0 mm. H, outer side of operculum (USNM 1020706). Scale = 200 µm. I, portion of radular ribbon (USNM 1020706). Scale = 20 µm. J, central radular teeth (USNM 1020706). Scale = 10 µm. K, lateral radular tooth (USNM 1020706). Scale = 10 µm.