Mycetobia christelae, Wojtoń & Kania & Krzemiński, 2019

Wojtoń, Maciej, Kania, Iwona & Krzemiński, Wiesław, 2019, Review of Mycetobia Meigen, 1818 (Diptera, Anisopodidae) in the Eocene ambers, Zootaxa 4544 (1), pp. 1-40 : 8-18

Mycetobia christelae sp. nov.

( Figs 10–13 )

urn:lsid:zoobank.org:act:F4116A01-D183-4F76-943F-AA4D862C6625

Diagnosis. Flagellomeres 1–6 short, distinctly wider than long, flagellomeres 7–13 longer than wide, up to 2x as long as wide; last flagellomeres longer than penultimate one, tapered at apex; last palpomere elongated, 3x as long as penultimate one, 3x as long as wide, rounded at apex, longer than second one; Sc ending before Rs bifurcation level; R 1 ending just before M 1+2 bifurcation level; R 2+3 elongated, 3x as long as Rs; the distance between R 1 and R 2+3 apices approximately 10x as long as the distance between Sc and R 1 apices; M 1+2 longer than M 1; fork of M 1+2 wide; the distance between M 1 and M 2 apices slightly longer than the distance between R 4+5 and M 1 apices; the distance between M 2 and M 3+4 apices as long as the distance between M 1 and M 2; R 2+3 ending before bifurcation of M 1+2 level, just beyond half R 4+5; dense setae on thorax, praescutellar and postalar setae not very elongated, pronotal setae thick and elongated; male terminalia large, approximately 0.25 the body length with cerci very elongated, massive, blunt ended and gonostyles shovel-shaped.

Etymology. The species name is dedicated to Christel Hoffeins from Hamburg, Germany, the owner of amber inclusions collection and the expert in Baltic amber inclusions.

Type material. Holotype: No. 1258-3 (male), Baltic amber, (coll. Ch. & H. W. Hoffeins); Paratype: No. 838—1 (male), Baltic amber, (coll. Ch. & H. W. Hoffeins); Additional material: No. MP /3694 (female), Baltic amber, ( ISEA PAS) .

Description. Body ( Fig. 11 A–F) 2.68–3.06 mm long. Head, thorax and legs brown. Head: antenna ( Figs 10A ; 12 A–C), 1.10 mm long, 14 flagellomeres with elongated sparse setae, shorter to width of segments bearing them; additionally very short setae on all flagellomeres; last flagellomere 0.11 mm long, penultimate one 0.03 mm; palpus maxillaris 4-segmented, the second palpomere 0.10 mm long, penultimate one 0.04 mm, last palpomere 0.14 mm long.

Thorax ( Fig. 12D, E ): prescutellar setae not very elongated, wing ( Figs 10C ; 12F ): 3.18 mm long, 1.06–1.14 mm wide; Sc ending opposite A 1 apex; R 1 apex in about 0.6 of wing length; Rs separate Rb in 0.4 of wing length from wing base; Rs 0.27 mm long; cross-vein r-m shorter than m-m.

Legs ( Figs 13E, F ): fore, middle and hind tibia with two unequal tibial spurs; tarsus comparatively elongated with last segment only slightly longer than penultimate one and not very elongated, rather slender claw and almost invisible empodium. Apical comb well visible on fore and hind legs; coxa of fore leg 0.62 mm, trochanter 0.13 mm, femur 0.86 mm, tibia 0.80 mm, tarsus 1.05 mm long (0.58/0.13/0.10/0.09/0.15). Tarsus of fore leg 0.3 the length of entire leg including coxa. Longer tibial spur of fore leg 0.07 mm long. Coxa of middle leg 0.60–0.70 mm, trochanter 0.10–0.12 mm, femur 0.77–0.93 mm, tibia 1.01–1.02 mm, tarsus 1.13–1.28 mm long (0.64–0.77/ 0.15–0.17/0.12–0.13/0.09–0.11/0.11–0.12). Coxa of hind leg 0.81 mm, trochanter 0.09–0.10 mm, femur 1.06–1.07 mm, tibia 1.10–1.22 mm, tarsus 1.05–1.28 mm long (0.59–0.77/0.11–0.19/0.11–0.12/0.09–0.10/0.11–0.13). All tibia with dense, not very elongated setae.

Abdomen ( Figs 11 A–F; 13A–D): male terminalia ( Figs 10B ; 13 B–D) very large, 0.79–0.80 mm long, dense, thick and elongated setae on anal point, gonocoxite, cerci and gonostyles, anal point wide, with many, dense setae, blunt ended; gonocoxite almost black with very elongated setae; female terminalia ( Fig. 13A ) comparatively massive.

Remarks: There are no differences in wing venation between males and females.

Comparison. M. christelae sp. nov. differ from other known from Baltic amber species by elongated last palpomere, which is 3x as long as penultimate one and is rounded at apex, not tapered as in other species of Mycetobia known from Eocene or described herein. This species also differ from other species in structure of male genitalia, e.g. in M. szwedoi sp. nov. additional structures are hooked, while in M. christelae they are shovelshaped. Moreover, M. christelae sp. nov. differs distinctly from M. silvia sp. nov. in shape of flagellomeres. In M. silvia sp. nov. flagellomeres are cylindrical, as long as wide up to twice longer than wide or even longer; in M. christelae sp. nov. flagellomeres 1–6 are much wider than long. Differences in wing venation are provided in Table 2.

Figures

FIGURE 10. Mycetobia christelae sp. nov., A. No. MP/3694 (female) (ISEA PAS), flagellomeres; B. holotype, No. 1258-3 (male), (coll. Ch. & H. W. Hoffeins), drawing of male terminalia; C. No. MP/3694 (female) (ISEA PAS), wing venation.

FIGURE 11. Mycetobia christelae sp. nov. A, B. holotype, No. 1258-3 (male) (coll. Ch. & H. W. Hoffeins). A. habitus, laterodorsal view; B. enlarged view of head, thorax and abdomen, latero-dorsal view; C., D. paratype, No. 838-1 (male) (coll. Ch. & H. W. Hoffeins); C. habitus, dorsal view; D. habitus, ventral view; E., F. No. MP/3694 (female) (ISEA PAS), habitus. E. lateral view; F. latero-ventral view.

FIGURE 12. Mycetobia christelae sp. nov. A.–C. flagellomeres: A. No. MP/3694 (female) (ISEA PAS); B. No. 838-1 (male) (coll. Ch. & H. W. Hoffeins), antenna; C. holotype, No. 1258-3 (male) (coll. Ch. & H. W. Hoffeins), enlarged view of flagellomeres; D. holotype, No. 838-1 (male) (coll. Ch. & H. W. Hoffeins), head and thorax, latero-ventral view; E., F. holotype, No. 1258-3 (male) (coll. Ch. & H. W. Hoffeins), E. thorax, lateral view; F. enlarged view of Sc, R 1 and R 2+3 apices. Remark and abbreviations: red points—the distance between red points is equal to Rs length.

FIGURE 13. Mycetobia christelae sp. nov. A. No. MP/3694 (female) (ISEA PAS), female terminalia; B, C. holotype, No. 1258-3 (male) (coll. Ch. & H. W. Hoffeins), male terminalia: B. lateral view; C. dorso-lateral view; D.–F. No. 838-1 (male) (coll. Ch. & H. W. Hoffeins): D. male terminalia, latero-ventral view; E. fore tibia with dense setae visible; F. middle and hind tibia with dense setae visible.