Helluodes Westwood

Sabu, Thomas K., Abhitha, Prabhakar & Zhao, Danyang, 2008, A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae), Zootaxa 1745 (1), pp. 30-46 : 35

Key to the species of Helluodes Westwood

1. Anterior margin of labrum with a distinct median emargination between rounded anterior angles ( Fig. 12 ); anterior angles of pronotum expanded, rounded and weakly acute; pronotum distinctly sinuated anterior to posterior angles; posterior angles of pronotum strongly reflexed and sharp at apex ( Fig. 14 ); ligula slen- der, widest at middle; TLA = 25–27.5 .................................................................... H. westwoodi Chaudoir.

- Anterior margin of labrum nearly straight or slightly emarginated between rounded anterior angles; anterior angles of pronotum expanded and rounded; pronotum weakly sinuated anterior to posterior angles; posterior angles of pronotum weakly reflexed and blunt at apex ( Figs. 8, 11 ); ligula slender and narrower towards apex................................................................................................................................................2

2 Anterior margin of labrum distinctly crenulate with 4 distinct indentations between anterior angles ( Fig. 6 ); maxillary palpomere 4 truncate and broadly dilated from middle to apex ( Fig. 7 ); TLA = 24–26.2 ...... .................................................................................................................................. H. devagiriensis n.sp.

- Anterior margin of labrum without crenulation and slightly emarginate between anterior angles ( Fig. 9 ); maxillary palpomere 4 roundly narrowed at apex and roundly dilated from middle to apex ( Fig. 10 ); TLA = 24.5 .................................................................................................................. H.taprobanae Westwood.

Figures

FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled.