Inca bonplandi ( Gyllenhal, 1817 )

Seidel, Matthias, Arriaga-Varela, Emmanuel & Sousa, Rafael, 2018, Catalogue of the Incini with the description of the first Archedinus species from Honduras (Coleoptera: Scarabaeidae: Cetoniinae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2), pp. 389-405 : 397-398

Inca bonplandi ( Gyllenhal, 1817)

Trichius bonplandi Gyllenhal, 1817: 196 [original combination]

Inca serricollis (partim): GORY & PERCHERON (1833): 108 [ I. bonplandi synonymized with I. serricollis ignoring priority]

Inca bonplandi: BURMEISTER & SCHAUM (1840) : 380 [new combination and resurrection of the senior synonym]

Ynca bonplandi: LAPORTE (1840) : 159 [combination with misspelled genus name]

Inca bomplandi: BRUCH (1911) : 216 [incorrect subsequent spelling, unavailable name]

Inca tapujo Perty, 1830 . Junior subjective synonym.

Inca tapujo Perty, 1830: 51 [original combination]. Synonymized with I. serricollis by GORY & PERCHERON (1833: 108).

Ynca tapago: LAPORTE (1840): 159 [combination with misspelled genus name and incorrect subsequent spelling of specific epithet, unavailable name]

Inca tapayo: GORY & PERCHERON (1833) : 108, 401 [incorrect subsequent spelling, unavailable name]

Inca bonplandi var. tapujo: SCHENKLING (1922) :4 [downgraded to variety, unjustified act]

Inca serricollis LePeletier & Serville, 1828 . Junior subjective synonym.

Goliath serricollis: DEJEAN (1821) : 61 [nomen nudum, unavailable name]

Inca serricollis LePeletier & Serville, 1828 [original combination].Synonymized with I. bonplandi and incorrectly treated as senior synonym by GORY & PERCHERON (1833: 108); priority corrected by BURMEISTER & SCHAUM (1840: 380).

Ynca servicollis: LAPORTE (1840):159 [combination with misspelled genus name; incorrect subsequent spelling, unavailable name]

Goliathus fasciatus Kirby, 1819 . Junior subjective synonym.

Goliathus fasciatus Kirby, 1819: 407 [original combination]. Synonymized by BURMEISTER & SCHAUM (1840: 380).

Inca fasciata: BLACKWELDER (1944) : 260 [new combination, incorrect gender agreement]

Goliathus inscriptus Kirby, 1819 . Junior subjective synonym.

Goliathus inscriptus Kirby, 1819: 407 [original combination]. Synonymized by BURMEISTER & SCHAUM (1840: 380).

Inca bonplandi var. inscriptus: SCHENKLING (1922) : 4 [downgraded to variety, unjustified]

Inca bonplandi v. inscripta: BLACKWELDER (1944) : 260 [incorrect gender agreement]

Goliathus tricuspis Drapiez, 1820 . Junior subjective synonym.

Goliathus tricuspis Drapiez, 1820: 272 [original combination]. Synonymized by BURMEISTER & SCHAUM (1840: 380).

Types. Inca bonplandi: One male syntype in NHRS and one female syntype in UUZM. Inca tapujo: Seven specimens of Inca bonplandi are deposited in ZSM and could be syntypes of I. tapujo ( SCHERER 1983) . Goliathus inscriptus: One male syntype in BMNH. Goliathus faciatus : one male syntype in BMNH. The types of Inca serricollis and Goliathus tricuspis were not located by us.

Locality records from literature. ARGENTINA: JUJUY ( BRUCH 1911), MISIONES ( BRUCH 1911, RICHTER 1913, DI IORIO 2013). BOLIVIA ( DI IORIO 2013). BRAZIL ( GYLLENHAL 1817; KIRBY 1819): BRASILIA (EVANGELISTA NETO et al. 2017), MINAS GERAIS ( PERTY 1830, PUKER et al. 2014), PARÁ ( SCHULZ 1901), RIO DE JANEIRO ( DRAPIEZ 1820, OHAUS 1900, COSTA et al. 1988, DI IORIO 2013). PARAGUAY: CAAGUAZÚ ( DI IORIO 2013), ITAPÚA ( DI IORIO 2013).

Additional material examined. ARGENTINA: MISIONES: 2 ♁♁ 1 ♀ ( NMPC): Argentina, Misiones, Iguazú, xii.99; 1 ♁ ( NMPC): Argentina, Misiones, Pozo Azul, xii.99; 1 ♁ 1 ♀ ( MSPC): Argentina, Misiones, San Pedro, vii.2010; 1 ♁ 1 ♀ ( ARPC): Argentina, Iguazu env.; 1 ♁ 1 ♀ ( BCRC):Argentina, Misiones, Mocona, iv.1985. BRAZIL: BRASILIA: 1♁ ( CEMT): Brazil, Distrito Federal, Fazenda Água Limpa, 15°57ʹ26.0″S, 47°56ʹ42.3″W,Armd.Isca Abacaxi 1, 28.xii.2013,Evangelista Neto, F.leg.; 1 ♀ ( MZSP): Distrito Federal, Floresta Cabeça, i.1957, B. Pereira leg.; 1 ♀ ( CEMT): Brazil, Distrito Federal, Fazenda Água Limpa, 15°57ʹ20.3″S, 47°56ʹ41.5″W, Armd. Isca Banana 12, 13.xi.2013, Evangelista Neto, J. ESPÍRITO SANTO: 1♁ 2♀♀ ( ZMUH):Brazil, Espirito Santo; 1♀ ( DZUP): Brazil, Espirito Santo, Linhares, (P. Sooretema), xii.1969, J. M. Lima leg. ( DZUP 273765). MINAS GERAIS: 1 ♀ ( NHRS): Minas Geraes, Mar d. Espanha, J. Zikan leg. PARANÁ: 2 ♀♀ ( NMPC): Paraná; 1 ♁ ( ZMUH): Parana, Curitiba; 1 ♀ ( CEMT): Brazil, Paraná, Londrina, 11.xii.1995, J. Lopes leg.; 1 ♀ ( ZMUH): Brazil, Parana, Lucena; 1 ♀ ( MZSP): Brasil, Rolandia, Norte-Paraná, iii.1951, Dirings leg. RIO DE JANEIRO: 1 ♁ 2 ♀♀ ( NMPC):Rio de Janeiro,Havlasa leg; 1♀ ( MZSP):Rio de Janeiro,Itatiaya, 700 m, 17.i.1926, J. F. Zikán leg.; 1 ♀ ( NMPC): Rio de Janeiro; 1 ♁ 1 ♀ ( NMPC): Petrópolis; 1 ♀ ( NHRS): Rio de Janeiro, Petropolis, 1.ii.1899, F. Ohaus; 1 ♁ ( NHRS): Rio de Janeiro, Corcovado, 9.i.1899, F. Ohaus; F. Ohaus; 1 ♁ ( NHRS): Rio de Janeiro, Corcovado, 9.i.1905, F. Ohaus; 1 ♀ ( NHRS): Rio de Janeiro, Petropolis, 2.ii.1899, F.Ohaus; 1♁ ( NHRS):Rio de Janeiro, Petropolis, 12.i.1899, F. Ohaus; 1 ♁ ( NHRS): Rio de Janeiro, Itatiaya, 27.xi.1926, F. Ohaus. RIO GRANDE DO SUL: 1 ♀ ( PSPC): Brazil, Rio Grande do Sul; 1 ♁ 1 ♀ ( NMPC): Rio Grande do Sul; 1 ♀ ( ARPC): Brasil,Rio Grande do Sul, Santa Cruz; 1♁ ( ZMUH):Brazil, P.Alegro; 1♁ 1 ♀ ( ZMUH): Rio Grande do Sul; 1 ♀ ( NHRS): Sao Leopoldo, J W. Stahl leg.; 1 ♁ ( FZBRS): Rio Grande do Sul, S. Luiz Gonzaga, 19.xii.1967, A. C. M. Moraes, (Col. MCN 163. 471); 1 ♁ ( FZBRS): Rio Grande do Sul, Santa Maria, (Cerrito), 11.ii.1994, L. Witeck leg.(Col MCN 218853); 1 ♀ ( MZUEFS): Brazil, Rio Grande do Sul, Vila Olivia, 20.ii.1950, J. Becker leg. (#30589); 1 ♁ ( MZUEFS): Brazil, Rio Grande do Sul, 14.ii.1950, J. Becker leg.(#30598). SANTA CATARINA: 1♀ ( NMPC):Santa Catarina.Fu[?] ke leg.; 1 ♀ ( MZSP): Brasil, Blumenau, Bairro Garoia, Moro Spitzkopf, iii.1962; 1 ♀ ( MZSP): Brasil, Santa Catarina, Joinville, Dirings leg.; 1 ♁ ( MZSP): Santa Catarina, Nova Teotonio, i.1949, F. Plaumann leg.; 1 ♀ ( MZSP):Santa Catarina,Rio Vermelho, i.1958, Dirings leg.; 1♀ ( BCRC): Brazil, Sao Bento do Sul; 1 ♀ ( ZMUH): Santa Catharina; 1 ♀ ( MZSP): Brazil, Santa Catarina, Timbó, viii.1963, Dirings leg. SÃO PAULO: 1 ♁ ( MZUEFS): Brazil, São Paulo, Aguas da Prata 27.ii.1969, J. Becker leg. (#30592);1♁ 1♀ ( NMPC):São Paulo,Mráz leg.;1♁ ( UNSM):Brazil,Sao Paulo,Itaici, i.1960,J.R.Neidoefer leg.; 1♀ ( MZSP):São Paulo,Ypiranga; 1 ♀ ( CMNC): Brazil, Sao Paulo, Capao Bonito, ii.1946, J. Guerin leg.; 1 ♀ ( ZMUH): Brazil, Sao Paulo, 1917; 1 ♀ ( ZMUH): Brazil, Sao Paulo; 1 ♀ ( MZSP): Brasil, São Paulo, Piracicaba, xi.1961, K. Reichardt leg.; 1 ♀ ( MZSP):São Paulo, São Sebastião, Praia Baraqueçaba, 2.iv.1963, P.Araujo leg.; 1 ♀ ( MZSP): Brazil, São Paulo, Sertãozinho, Res. Ecol. Augusto Ruschi, 21°10.520’S, 48°5.47’W, 529 m, 07–21.xii.2011. Malaise borda, V. C. Silva, P. F. Donda & G. P. Ignácio leg. PARAGUAY: GUAIRÁ: 1 ♁ ( NMPC):Paraguay,Villarica,Kolonio Sudetica. ITAPÚA: 2♁♁ 1♀ ( ARPC): Paraguay, Encarnacion, xii.1975. ITAQUIRI: 1♀ ( DZUP):Paraguai, Itaquiri, 400 m, 15–20.i.1980, Mielke & Miers leg. ( DZUP 273741). SAN PEDRO: 1 ♁ 2 ♀♀ ( NMPC): Paraguay, Carlos Pfannl, i.1969, José Andres lgt.

Remarks. BLACKWELDER (1944) erroneously attributed authorship of Inca serricollis to GORY & PERCHERON (1833). SCHENKLING (1922) listed “ tapujo ” and “ inscriptus ” as variations of Inca bonplandi (Gyllenhall, 1817) , and there seems to be no evident justification for Schenkling’s listing. KRAJCIK (1999) incorrectly gave the year for Inca tapujo as 1834 and not 1830. BURMEISTER & SCHAUM (1840) stated that Goliathus inscriptus Kirby, 1819 only represents an “insignificant variety” and later BURMEISTER & SCHAUM (1841) added that it is only a darker variant of Inca bonplandi . Goliathus fasciatus Kirby, 1819 and Goliathus tricuspis Drapiez, 1820 were listed by BURMEISTER & SCHAUM (1840) as synonyms of Inca bonplandi without further comment. DI IORIO (2013) interpreted the synonymical listing of Goliath fasciatus and Goliath inscriptus under Inca bonplandi only as a reference, but we acknowledge that BURMEISTER & SCHAUM (1840) established these synonymies. SCHAUM (1849) stated that the type of Goliathus fasciatus Kirby, 1819 only represents a “slight variation” of Inca bonplandi . He clearly intended to keep Kirby’s species in synonymy. BLACKWELDER (1944) unjustifiably emended and combined the name to Inca fasciata . Both, BURMEISTER & SCHAUM (1840) and LAPORTE (1840) resurrected Inca bonplandi as valid name over the junior synonyms, but BURMEISTER & SCHAUM (1840) was published in July–September 1840 ( BOUSQUET 2012), whereas LAPORTE (1840) was published as earliest as on 26 December 1840 ( BOUSQUET 2016). Therefore, BURMEISTER & SCHAUM (1840) deserve credit for the resurrection.