Ulidia omani Steyskal

Morgulis, E. & Freidberg, A., 2014, The Ulidiini (Diptera: Tephritoidea: Ulidiidae) of Israel, with a key to the world species of Ulidia and description of five new species, Zootaxa 3780 (2), pp. 201-247 : 229-230

Ulidia omani Steyskal

( Figs. 25 , 37 , 59 , 66 , 82 , 96 , 107 )

Ulidia omani Steyskal, 1970: 227

Material examined. ISRAEL: Type material: Paratype: Be`er Sheva' [Beersheba], 24.iii.1964 [64], P. Oman (1♂, 2♀) ( USNM); Non-type material: Jordan Valley and Southern Golan: Mehola [Mechola], 26.ii.1972, J. Kugler (1♀); Nahal Tirza [W. Faria], 4.iii.1985, I. Susman (1♂); Judean Desert: Yeriho [Jericho], 20.ii.1971, J. Kugler (4♂); Midbar Hevron [Hebron Desert], 26.iii.1974, D. Furth (10♂, 4♀); Northern Negev: Nahal Besor [N. Bsor], 5.iii.1970, J. Kugler (2♂, 2♀); 'En Besor [Ein-Bsor], 31.iii.1975, F. Kaplan (1♂); 'Arad, 570 m, 31º15.346'N 35º11.958'E, 7.iii.2010, E. Morgulis (2♂, 1♀); Nahal Ye'elim [Ye'elim nr. Arad], 8.iv.1998, A. Freidberg (1♀); Nahal Sekher [Nahal Sekher], 6.iv.1992, A. Freidberg (32♂, 34♀); Mishor Rotem [Mishor Rotem], 9.iii.1965, M. Weichselfish (1♀); Be`er Sheva' [ Beer Sheba], 24.iii.1954, O. Theodor (1♂, 1♀); Be`er Sheva'-Dimona [ Beer Sheva Dimona], 10.iii.1965, M. Weichselfish (2♂, 2♀); ze'elim [Zeelim], 12.iii.1974, D. Furth (2♂, 2♀); Dimona, 19.iii.1978, D. Furth (1♀), 28.iii.1978 (1♂); 12.iv.1990, A. Freidberg (6♂); Bor Mashash, 25.iii.1987, A. Freidberg (2♂, 2♀), F. Kaplan (5♂, 1♀), 16.iii.1995, A. Freidberg (1♀); Bor Mashash [Be`er Meshash], 20.iii.1977, A. Freidberg (1♂); Dead Sea Area: Nahal Zohar Spill [Shefech Zohar], 8.iii.1972, J. Kugler (1♀); Central Negev: Mash'abbe Sade, 31.iii.1975, F. Kaplan (1♀), 19.iii.1978, A. Freidberg (1♂), 12.iii.1981, D. Furth (1♀), 31.iii.1981, F. Kaplan (1♂); Ma'agar Yeroham, 30º59'25.8''N 34º53'47.2''E, 8.iii.2010, A. Freidberg (6♂, 10♀), E. Morgulis (22♂, 24♀); Park Yeroham [Park Yeroham], 7.iv.1998, A. Freidberg (1♂); Yeroham [Jeruham], 21.iii.1971, J. Kugler (2♀); Yeroham [Yeroham], 22.iii.1971, J. Kugler (9♂, 7♀), 25.iii.1987, F. Kaplan and A. Freidberg (1♀, 1♂), 20.iv.1992, A. Freidberg (1♂); Horbat Shivta [Shivta], 13.iii.1977, A. Freidberg (1♂); Sede Boqer, 28.iii.1978, D. Furth (1♂); 'En Mor, 30º50.676'N 34º46.603'E, 18.iii.2010, A. Freidberg (26♂, 6♀); 'En Mor [Ein Mur], 16.iii.1970, J. Kugler (1♀), 31.iii.1981, A. Freidberg (1♂, 1♀), F. Kaplan (1 ♂); 'En Mor, Nahal Hawarim, 30º50.676'N 34º46.603'E, 4.iv.2010, E. Morgulis (1♀); 'En 'Avedat [E. Avdat], 9.iii.1973, D. Furth (1♀); 'En 'Avedat [Ein Avdat], 29.iii.1980, W. N. Mathis and A. Freidberg (17♂, 11♀); Mizpe Ramon [Mitzpe], 2.iii.1979, D. Furth (1♂); Mizpe Ramon [Mizpe Ramon], 6.iv.1992, A. Freidberg (1♂); Mizpe Ramon, 5km N [Mitzpe Ramon 5km N], 17.iii.1995, A. Freidberg (1♂); Mizpe Ramon, observatory [Mizpe Ramon observatory], 17.iii.1995, A. Freidberg (1♂); Nahal Loz, Har Ramon [Nahal Loz Har Ramon], 900 m, 14.iv.1992, A. Freidberg (2♂, 1♀); Har Hemet, near Makhtesh Ramon, 14.iv.1992, A. Freidberg (1♂, 2♀); Har Horesha [har horesha], 17.iii.1995, A. Freidberg (1♀)( TAUI).

Description. Head ( Fig. 25 ): Structure: 0.86–1.05 times as high as long. Frons of same width posteriorly and at lunule level, 0.94–0.97 times as wide as long, smooth. Eye 0.89–1.01 times as high as long. Gena in male 0.37– 0.40 times as high as eye, in female 0.47–0.59 times as high as eye; in both sexes gena posteriorly strigose. Frontoorbital plate at antenna insertion level 0.28–0.38 times as long as eye, smooth. Color and vestiture: Head brown to black. First flagellomere brown, silvery microtrichose. Arista with microscopic rays. Setae and setulae: Medial vertical seta as long as lateral vertical seta; ocellar seta 0.42–0.54 times as long as medial vertical seta; postocellar seta missing in 70% of specimens, weakly developed in 20% of specimens and well developed (as long as ocellar seta) in 10% of specimens; orbital seta 0.48–0.70 times as long as medial vertical seta. Frons short setulose. Vibrissal angle bare. Gena and postgena with fine, short setulae; 1 short and thin genal seta. Palpus and labellum with short black setulae.

Thorax: Color and vestiture: Subshiny black, densely shagreened. Setae and setulae: 2 supra-alar, 1 dorsocentral, 1–3 anepisternal and one katepisternal setae present; 1 acrostichal setula present. Postpronotal lobe, scutum, scutellum, anepisternum and katepisternum, setulose; setulae 0.20–0.25 times as long as major setae.

Legs: Entirely blackish-brown. Hindfemur not widened, as wide as midfemur in both sexes.

Wing ( Fig. 37 ): Pattern: wing with basal-cubital cell and basal half of cubital cell brown; hyaline area from level of mid-cubital cell to level of crossvein bcu, infuscate apical to level of crossvein bcu; pterostigma mat brown. Venation: Veins yellowish-brown to black. Crossvein R-M aligned apical to apex of vein R1. Ratios: Cell r4+5 0.20–0.25; vein Cu2 0.69–0.83. Calypteres white. Halter base and stem brown, knob yellow.

Abdomen: Tergites strongly shagreened and setulose.

Male terminalia: Epandrium ( Fig. 59 ) 1.51–1.52 times as high as wide; cerci slightly separate, truncate, not triangular; lateral surstylus apically bilobate, posterior lobe medially inclined; medial surstylus with 2 prensisetae and 10–15 setulae in 3 rows. Phallus ( Fig. 66 ) with 1 caecum. Glans ( Fig. 82 ) rounded, with large apical elongations.

Female terminalia: Aculeus at widest point 7.8–9.0 times as long as wide. Cercal unit ( Fig. 96 ) triangular, apically pointed. Sensilla: BV slightly basal to BD. Spermathecae ( Fig. 107 ) globulose with smooth surface.

Measurements (mm): Body length 2.6–3.6, wing length 2.3–2.8.

Type locality. Beersheba (=Be`er Sheva'), Israel.

Distribution. Israel.

Adult activity in Israel: End of February to middle of April.

Diagnosis. This species differs from its congeners in the following combination of characters: densely shagreened thorax, absence of dorsocentral setae, and infuscate wing.

Comments. This species resembles U. fascialis Hendel, 1931 . Two paratypes (♀) of U. fascialis were examined and no external differences between the two species were found. Examination of the male terminalia is required in order to decide whether these are distinct species or U. omani is a junior synonym of U. fascialis Hendel.

The original spelling of the species name is " fascialis "; the spelling " facialis " ( Hennig 1940; Kameneva 2008) is an unjustified emendation.

Figures

FIGURES 23 – 28. Head, lateral view. 23. Ulidia hirsuta n. sp.; 24. U. nigripennis; 25. U. omani; 26. U. ruficeps; 27. U. wadicola; 28. U. wasimi n. sp.

FIGURES 29 – 38. Wing. 29. Physiphora alceae; 30. Timia anomala; 31. T. sp. cf. berlandi; 32. T. ritae n. sp.; 33. Ulidia albidipennis; 34. U. aurata n. sp.; 35. U. hirsuta n. sp.; 36. U. nigripennis; 37. U. omani; 38. U. wasimi n. sp.

FIGURES 46 – 62. Epandrium, posterior view. 46. Physiphora alceae; 47. P. smaragdina; 48. Timia anomala; 49. T. sp. cf. berlandi; 50. T. fallax n. sp.; 51. T. jakowlewi; 52. T. libani; 53. T. ritae n. sp.; 54. T. xanthaspis; 55. Ulidia albidipennis; 56. U. aurata n. sp.; 57. U. hirsuta n. sp.; 58. U. nigripennis; 59. U. omani; 60. U. ruficeps; 61. U. wadicola; 62. U. wasimi n. sp.

FIGURES 63 – 68. Coeca and setae of the phallus. 63. Timia anomala; 64. T. fallax n. sp.; 65. Ulidia albidipennis; 66. U. omani; 67. U. parallela; 68. U. wasimi n. sp.

FIGURES 78 – 85. Glans. 78. Ulidia albidipennis; 79. U. aurata n. sp.; 80. U. hirsuta n. sp.; 81. U. nigripennis; 82. U. omani; 83. U. ruficeps; 84. U. wadicola; 85. U. wasimi n. sp.

FIGURES 93 – 99. Cercal unit, ventral view. 93. Ulidia aurata n. sp.; 94. U. hirsuta n. sp.; 95. U. nigripennis; 96. U. omani; 97. U. ruficeps; 98. U. wadicola; 99. U. wasimi n. sp.

FIGURES 100 – 109. Spermatheca. 100. Physiphora alceae; 101. P. smaragdina; 102. Timia anomala; 103. T. ritae n. sp.; 104. Ulidia albidipennis; 105. U. aurata n. sp.; 106. U. hirsuta n. sp.; 107. U. omani; 108. U. ruficeps; 109. U. wasimi n. sp.