Kmentiini, Stehlík, 2013

Stehlík, Jaroslav L., 2013, Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae), Acta Entomologica Musei Nationalis Pragae 53 (2), pp. 505-584 : 565-566

Kmentiini trib. nov.

Type genus. Kmentia gen. nov., here designated.

Diagnosis. Labium reaching between mesocoxae. Callar lobe weakly gibbose in both sexes. Ventral surface of profemora in both sexes with only a short longitudinal furrow limited to its apical third; basal two-thirds medially convex with numerous denticles on its entire surface ( Figs 83 , 97 ). Protibia in both sexes unarmed ( Figs 95−96 ). Parameres horizontally positioned, their apices approaching and standing opposite each other ( Fig. 85 ), bent dorsally in their apical portion, apex hook-shaped, pointed dorsally ( Figs 85, 87 ), distinctly surpassing ventral rim of pygophore (see in lateral view − Fig. 88 ).

Differential diagnosis. The tribe Kmentiini trib. nov. Belongs in the subfamily Physopeltinae , sharing its important synapomorphy – the intersegmental sulci between abdominal sternites deeply sinuated (S-shaped). It differs from the nominotypical tribe Physopeltini in the characters listed above, especially in the structure of profemora and male genitalia.

Genus included. The tribe is currently monotypical, including only Kmentia gen. nov.

Discussion. The West African Physopelta festiva ( Fabricius, 1803) is excluded from Physopelta based on a number of differences in its external morphology and structure of genitalia. It is interesting that some of these characters resemble the New World subfamily Larginae , especially in the structure of the profemora (lacking the longitudinal furrow ventrally) and parameres. Although the parameres of all the remaining Physopeltinae are apically obtuse, directed skewed towards the center of the pygophore, and do not surpass the ventral rim; the parameres in Physopelta festiva are long, parallel, surpass the ventral rim, and have hookshaped apices, which is characteristic for Larginae . In addition, the flat callar lobe of both sexes distinguishes Ph. festiva from all the remaining Physopelta species. Considering the differences between Ph. festiva and the remaining Physopeltinae taxa, I establish a new tribe, Kmentiini trib. nov., and a new genus, Kmentia gen. nov.

Figures

Figs 81–84.Kmentia festiva (Fabricius, 1803), male.81 – fore coxae and trochanters (magnification 47×); 82 – detail of procoxa and protrochanter (70×); 83 – ventral surface of profemur (32×); 84 – detail of peritreme (150×). Scale bars: 0.2 mm (Fig. 84), 0.5 mm (Figs 81–83). (SEM micrographs: P. Kment).

Figs 93–98. Kmentia festiva (Fabricius, 1803). 93 – female; 94 – male; 95–96 – fore leg (95 – large male, 96 – small male); 97 – ventral surface of profemur. 98 – Physopelta (Neophysopelta) gutta gutta (Burmeister, 1834), ventral surface of profemur. (Photos: L. Dembický).

Figs 85–92. Kmentia festiva (Fabricius, 1803), male. 85–88 – pygophore: 85 – intact, dorso-posterior view (magnification 50×), 86 – dissected, dorsal view (55×), 87 – intact, posterior view (50×), 88 – intact, lateral view (60×). 89–92 – paramere (different orientations; 150×). Scale bars: 0.2 mm (Figs 89–92), 0.5 mm (Figs 85–88). (SEM micrographs: P. Kment).