Helluonini, Hope, 1838

Raj, Shiju T., Sabu, Thomas K. & Danyang, Zhao, 2012, The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns, Insecta Mundi 2012 (206), pp. 1-15 : 2

Key to the Indian genera of the tribe Helluonini (modified from Sloane 1914)

1. Wingless forms; mentum with a strongly developed stout median tooth; legs stout. ................. ......................................................................................................................... Omphra Dejean

– Winged forms; mentum with an elongated spiniform median tooth; legs slender. ................... 2

2(1). Tarsi with fourth joint deeply bilobed. ............................................................ Creagris Nietner

– Tarsi with fourth joint entire. ...................................................................................................... 3

3(2). Fourth maxillary palpomere securiform. ....................................................... Colfax Andrewes

– Fourth maxillary palpomere not securiform. ............................................ Macrocheilus Hope