Syspasis haesitator (WESMAEL, 1845)

Riedel, Matthias, 2021, Revision of the Palaearctic species of Syspasis TOWNES, 1965 (Hymenoptera, Ichneumonidae, Ichneumoninae), Linzer biologische Beiträge 53 (2), pp. 775-810 : 782-784

Syspasis haesitator (WESMAEL, 1845) (figs 2, 18, 26, 36, 50)

Original: Ichneumon haesitator WESMAEL, 1845: 36 .

L e c t o t y p e: (♀) 119; Monographie exempl. décrit; Ichneumon haesitator ♀ dét. C. Wesmael; Lectotype Ichneumon haesitator WES. Tow. ´65 (Brussels), type studied.

syn. Ichneumon operosus BERTHOUMIEU, 1901: 320, type lost, synonymized by DILLER & HORSTMANN (1997).

M a t e r i a l e x a m i n e d: 10♀♀ and 3♁♁ from Austria, Belgium, Finland, Georgia, Germany, Sweden, Russia Far East.

D e s c r i p t i o n: ♀: Body length 8-10 mm. Flagellum with 28-30 flagellomeres, moderately lanceolate; 1 st flagellomere 2.1-2.4× and 2 nd flagellomere 2.0-2.1× longer than wide; 7 th flagellomeres square; widest flagellomeres 1.5× wider than long. Temple moderately and roundly narrowed behind eye. OED 1.3-2.0× ocellar diameter. Frons transversely rugose-punctate. Face punctate and granulate. Clypeus slightly convex, punctate. Gena densely punctate ventrally, 0.85-0.9× as wide as eye. MI 0.9-1.1. Mesosoma with short pale brownish hairs. Side of pronotum punctate dorsally, striate ventrally. Notaulus indistinctly impressed. Lateral field of mesoscutum densely and superficially punctate and granulate. Mesopleuron punctate, partly striate. Metapleuron punctate. Scutellum wider than long, with scattered punctures, shining, without lateral carina. Horizontal part of propodeum rather long. Area basalis without distinct median tubercle. Area superomedia as long as or slightly longer than wide, hexagonal or halfmoon-shaped, finely rugose; anterior transverse carina indistinct. Area externa punctate and granulate. Area petiolaris transversely rugose, without lateral carina. Hind coxa densely punctate and granulate, without scopa. Hind femur densely punctate, 3.5-3.7× longer than wide. Hind tibia with 2-3 denticular spurs apico-externally. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 2-3× their width; vein 2m-cu reaching its middle. Vein 1cu-a slightly postfurcal. Metasoma oxypygous. Lateral field of postpetiolus c. 0.5× as wide as median field, the latter finely rugose or aciculate. Gastrocoelus indistinctly impressed. Thyridium transverse, 1.5-2.0× wider than the interval between thyridia. 2 nd tergite 0.70-0.77× as long as wide. 2 nd and 3 rd tergites superficially punctate, finely granulate and weakly striate medially. Ovipositor sheath moderately extending behind metasomal apex (by 0.3-0.7× length of 7 th tergite).

Color: black. Stripes or rings on flagellomeres 5/7-11, narrow frontal orbit, and often collar ivory. Vertex sometimes paler but without distinct ivory spot. Scape reddish ventrally. Apical half of postpetiolus and 2 nd to 4 th tergites red (4 th tergite sometimes black in apical half); tergites otherwise black. Coxae and trochanters black; legs otherwise red; hind femur mainly black; hind tibia apcially and hind tarsus ± blackish. Pterostigma brownish.

♁: Body length 9.5-10.0 mm. Flagellum with 29 flagellomeres, bristle-shaped; 1 st flagellomere 2.5× longer than wide. Tyloids on flagellomeres 6/7-12, long-oval, centrally situated, maximally 0.6× as long as their flagellomeres. Temple moderately and roundly narrowed behind eye. OED 1.4× ocellar diameter. Frons punctate and granulate dorsally, smooth above antennal sockets. Face densely punctate and granulate, dull. Clypeus densely punctate, apical margin sharp, straight. Gena with dense punctures. Malar space 0.6× as long as width of mandibular base. Mandible with two teeth, lower tooth small and strongly bent inwards, mandible appearing almost sickle-shaped. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma covered with dense grey hairs. Side of pronotum with very superficial punctures dorsally and with longitudinal rugae ventrally and caudally. Notaulus weakly impressed frontally. Mesoscutum with dense punctures and distinct granulation, dull. Mesopleuron densely punctate, partly rugose; speculum partly smooth and shining. Metapleuron punctate, shining. Juxtacoxal carina present. Scutellum moderately elevated, 1.3× wider than long, punctate, with lateral carina in basal 0.3. Area basalis with distinct median tubercle. Area superomedia heart-shaped, slightly wider than long; anterior transverse carina distinct, in its middle. Hind femur densely punctate, 3.6× longer than wide. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 3× their width; vein 2m-cu in its middle. Vein 1cu-a almost interstitial. Postpetiolus moderately widened, without dorsal carina, but median field moderately elevated. Lateral field punctate, median field with fine aciculation. Gastrocoelus narrow. Thyridium large, slightly oblique, 2.0× wider than the interval between thyridia. 2 nd tergite 0.85× as long as wide. 2 nd and 3 rd tergites coarsely but superficially punctate, granulate, dull. 4 th tergite with very superficial punctures, ± shining.

Color: black. Palps, often lateral spot on clypeus, facial and frontal orbits, often facial spots below antennal sockets, narrow stripe on vertex, sometimes also on outer orbit and on ventral scape, collar, sometimes hind edge of pronotum, spot on subtegular ridge ivory. Tergites black, sometimes with narrow apical dark reddish shine. Coxae, trochanters and trochantelli black; fore and mid legs otherwise red; hind femur black; hind tibia reddish, infuscate in apical 0.2-0.3; mid and hind tarsi brown to black. Wings hyaline; pterostigma brown.

Host: Panolis flammea (DENIS & SCHIFFERMÜLLER, 1775) (Lep., Noctuidae ) ( SACHTLEBEN 1929).