Archierato panamaensis ( Carpenter, 1856 ), 2020

Fehse, Dirk & Simone, Luiz Ricardo L., 2020, Contributions to the knowledge of the Eratoidae. X. Revision of the genus Archierato Schilder, 1933 (Mollusca: Gastropoda), Zootaxa 4851 (1), pp. 81-110 : 86-87

Archierato panamaensis ( Carpenter, 1856)

( Figs. 9 , 18 , 44 )

Erato? maugeriae var. panamaensis Carpenter, 1856: 162 ; Keen, 1968: 435 (pl. 58, figs. 66a, 66b); Cate, 1977: 361 (text fig. 45).

Erato panamaensis: Smith, 1910: 14 .

Erato (Hespererato) panamaensis: Keen, 1971: 488 (fig. 914).

Lectotype. NHMUK 1966619 /1; Hugh Cuming collection; designation by Keen (1968: 435). L 7.5 mm, W 5.2 mm, D 3.5 mm, CT –, LT 14

Type locality. ‘Pacific Panama’.

Etymology. Named after the type locality.

Diagnosis. Shell large, 7 to 9 mm in length, robust, pyriform, inflated, with widely spaced, coarse labral and posteriorly obscured columellar dentition; ventral fold thick, 16 labral teeth; maximum globosity at posterior third.

Description. Shell large, inflated, pear-shaped, with slightly elevated spire. Protoconch and subsequent whorls covered by thin callus. Suture distinct. Body whorl ~90% of total height, maximum diameter at posterior third, convexly tapered anteriorly. Anterior ventral margin largely indented. Dorsum rounded. Dorsal sulcus absent. Whole shell surface covered by very thin, sub-glossy callus. Aperture ~90% of total height, sinuous and slightly widened. Labrum narrow, thickened, smooth, flattened ventrally. Outer labral margin rounded, callused, edged at inner margin. Labral teeth coarse, widely spaced, irregular, 14 in number, restricted to labral edge. Siphonal canal extended, rounded, indented. Anal canal simple, blunt. Columella sinuous, curved without inner carinal ridge and parietal lip. Columellar denticles few, fine, posteriorly obscured, anterior most thickened forming terminal ridge. Fossula obscured, not delimited from columella. Callosities white; anterior tip very pale white-brown; dorsum brown.

Variability. Not enough specimens are available to evaluate intraspecific variability.

Distribution. Carlsbad, California, U.S.A., to Panama. ‘Bahía Honda, Veraguas, Panama’ ( Vega & González, 2002: 30). Confirmed by a specimen in the author’s collection: Carlsbad, California, U.S.A. (DFB 12089).

Material examined. W. Panama: Gobernadora Island, Gulf of Montijo ( DFB 5604 ) . U.S.A. California; Carlsbad , ( DFB 12089) .

Remarks. The shell of Archierato panamaensis is not as colourful as that of A. columbella —only the dorsum is coloured. It is distinguished from the latter species by the inflated shell, the wider, sinuous aperture, the narrow, non-declivous labrum, widely spaced, lesser developed labral denticles, the obscured columellar denticles, and the indented anterior tip (compare Figs. 39 and 40 ).

Figures

FIGURES 7–9. Archierato type specimens. 7. Archierato maugeriae (Gray in Sowerby I, 1832) lectotype NHMUK 197410/1, L 5.2 mm “Levy County, Florida, U.S.A.” (courtesy H. Taylor). 8. Archierato columbella (Menke, 1847) neotype LACM 3302, L 6.8 mm, White Port, Los Angeles County, California, USA (courtesy L.T. Groves). 9. Archierato panamaensis (Carpenter, 1856) lectotype NHMUK 1966619/1, L 7.5 mm, ‘Pacific Panama’ (courtesy H. Taylor).

FIGURES 16–18. Archierato specimens. 16. Archierato columbella (Menke, 1847) DFB 5603-3, L 5.6 mm, Encinitas, California, USA. 17. A. columbella DFB 2883-2, L 7.3 mm, Carlsbad, California, USA. 18. Archierato cf. panamaensis (Carpenter, 1856), DFB 5604-1, L 7.5 mm, Gobernadora Is, Gulf of Montijo, W Panama.

FIGURES 42–46. Comparison of the E Pacific Archierato species (the shells are equally sized for better comparison, consider, e.g., Tab. 1 for their real dimension). 42. A. michaelmonti, holotype. 43. A. rhondae, holotype. 44. A. panamaensis, lectotype. 45. A. columbella, neotype. 46. A. galapagensis, holotype.

FIGURES 36–41. Comparison of the W Atlantic Archierato species (the shells are equally sized for better comparison, consider, e.g., Tab. 1 for their real dimension). 36. A. guadeloupensis, holotype. 37. A. dalli, holotype. 38. A. dalli, off Florianópolis, Santa Catarina State, Brazil. 39. A. martinicensis, holotype. 40. A. maugeriae, lectotype. 41. A. janae, holotype.