Tracholena paniense, Dugdale, John S., 2005

Dugdale, John S., 2005, Three new species of Tracholena Common, 1965 (Lepidoptera: Tortricidae: Tortricinae: Schoenotenini) from New Caledonia associated with Araucariaceae, Zootaxa 870, pp. 1-16 : 9-11

Tracholena paniense sp.nov. ( Figs. 3 , 4, 7 , 11, 14, 15 )

Material examined: Holotype: Male, “NOUVELLE CALEDONIE: Mt Panié 1380 m, to light, 11 Dec.1990, J.S. Dugdale.” “ HOLOTYPE male/ Tracholena paniense Dugdale ” [red card], MNHN.

Paratypes. New Caledonia: 4 males, 6 females, same data as Holotype except capture dates span 4–15 December 1990. 1 male, one female, ONNC; 3 males, 5 females. NZAC.

Description: Male ( Fig. 3 ), female: Head with vertex tufts pale with pale­ brown tips; antennal scape pale brown, flagellomeres with pale­brown and scattered black scales, flagellomeres 1–10 cylindrical, 11ultimate each tapered apically; each flagellomere ( Fig. 7 ) with a basal whorl of sinuous trichomes 2.0X flagellomere width; paired sensilla coeloconica subapical and sub­lateral on each flagellomere, separated by at least their diameter. Labial palp second segment buff basally, pale brown apically, vom Raths organ less than half the length of apical segment. Maxillary palp 2­segmented on pedicel, basal segment broadly fused with pedicel and with 2 mesal setae, apical segment basally narrowed, roughly diamond shaped, and with 4 scales irregularly arranged. Prosternal midline scales pale buff contrasting with the brown and blackish speckled fore­coxa. Forewing: As in Fig. 4 , length 2.4X width (males), narrower,>2.6X (females); forewing rusty­ochreous brown (concolouus with head and mesonotum); forewing colour pattern: basal area with a short longitudinal line of black scales along vein R; ante­median line from costa to dorsum with middle third absent, marked by raised black scales bordered by orange and buff scales; median area with 2 irregular pale areas towards costa and dorsum respectively; post­median line arising costally in a dark scale patch, absent discally, marked from vein CuA2 as for ante­median line, and forming the inner border of a conspicuous dark “thumbprint” present in some specimens; post­median area to termen with transverse irregular strigae of orange scales mixed with black scales; a conspicuous triangular patch costally, pre­apically. Hindwing: Shining buff (pale grey in liparodes ); anal tuft length about half that of anal margin. Wing venation ( Fig. 4 ): Veins M2, M3, and Cu1A separate at base ( cf. liparodes , Fig. 6 ).

Wingspan: Holotype male: 17.8 mm; 4 paratype males: 16.0– 17.6 mm; 6 paratype females: 14.4–17.6 mm.

Male genitalia ( Fig. 11 ): Uncus slender, curved, tubular; uncus­tegumen junction transverse, straight; socii large, lobate, pendent, with a file of erect setae outwardly near base; gnathos arms forming a narrow V, smooth, apices fused, apical hook very short, apex truncate. Transtilla broadly W­shaped, the two triangular, apically blunt­spinose processes directed postero­ventrally; juxta apically truncate, weakly keeled along mid­line. Valva wide, with costa concave and sclerotised to valval apex. Aedeagus cylindrical, caecum penis very long (bulbus ejaculatorius entering just before aedeagal apex), apex blunt, unadorned; bulbus ejaculatorius tube with about 20 spirals.

Female genitalia ( Figs. 14, 15 ): Ovipositor lobes slender, elongate, parallel and with anterior apices lobate, deflected mesally in dry specimens; ovipore level with anterior third of lobes, with a V­shaped mesal lobe; sternite 8 with a transverse scobinate ridge; lamella post vaginalis a broad and deep scobinate field with an anterior sclerotised strip; lamella ante­vaginalis a narrow spinulose field; ostiolar sclerite present, extending beyond ductus which arises at half­sclerite length; ductus seminalis arising just anterior and ventral to the colliculum; ductus bursae and cestum with 20 spirals, slender, elongate; corpus bursae ovoid, smaller than bulla seminalis, signum lacking.

Diagnosis: This relatively broad­winged moth (forewing length <2.75X width) differs from the other broadwinged Tracholena species ( T. homopolia , T. lipara and T. liparodes ). In T. homopolia the forewing scales are not uniformly grey with white tips; the gnathos arms lack spinules; the apices of the transtilla are deflected; apical spinules are absent on the aedeagus; and there are 20, not 12 spirals in the helical ductus bursae (cf. Fig. 14 and Common 1965: figs. 15K, L, and 18B). In T. lipara the forwing ground colour is ochreous, not grey; the hindwings are paler; the flagellomere trichomes are longer than the flagellomere and arranged in a basal fascicle; the gnathos arms are smooth; the transtillar spining is confined to the apex of each transtilla process; the entry of the bulbus ejaculatorius is subapical; and there are more spirals in the ductus bursae (20, not 14) (cf. Fig.14 and Common 1973: figs. 1–3). It can be distinguished from T. liparodes by the ochreous brown forewing ground colour (grey in liparodes ) and pale whitish or buff hindwing (pale grey in liparodes ); the longer flagellomere basal trichomes; the well­separated sensilla coeloconica (contiguous in liparodes ); the widely separated bases of hindwing veins M2, M3, and Cu1A (M3, Cu1A stalked in liparodes ); the smooth gnathos arms (spinulose in liparodes ), the deflected transtilla arms with spines confined to the apex of each lobe; the aedeagus lacking a hooked process and cornuti (both present in l iparodes); the ovipositor lobes with recurved anterior apices (straight in liparodes ); the helical cestum/ductus bursae with 20 spiral twists; and the corpus bursae lacking a signum (cestum with one extended spiral and signum present in liparodes ).

General notes: The type series was collected at light under a canopy of Agathis mon ­ tana (with scattered peripheral Araucaria spp.) on the shoulder below the summit block of Mt Panié between 4 and 15 December 1990. No damage ascribable to this species was found on Agathis ; the few Araucaria male cones I could find were unblemished, but were too few to base any conclusion on.

Etymology: The name is derived from the locality, Mt Panié.

Figures

FIGURES 1 – 3. Fig. 1. Tracholena nigrilinea sp. nov. Male, Ponerihouen, habitus. (LH wings omitted). Fig. 2. T. liparodes sp. nov. PT male, Mt Panié, habitus. Fig. 3. T. paniense sp. nov. PT male, Mt Panié, habitus. (Scales represent 0.5 mm)

FIGURES 1 – 10. Fig. 4. T. paniense. PT male fore and hind wing venation [NZAC prep. 875]. Fig. 5. T. nigrilinea. Fore and hind wing venation, male [NZAC prep. L 862]. (Outer scale tuft across vein CuA 2 omitted.) Fig. 6. T. liparodes sp. nov. Hind wing M 2 ­ M 3 ­ CuA 1 disposition. Fig. 7. T. paniense, male flagellomere 10, ventral, showing widely separated sensillae coeloconica. [NZAC prep. L 879]. Fig. 8. T. nigrilinea, PT male, genitalia, Ponerihouen [NZAC prep. L 859]. Fig. 9. T. liparodes, PT male, aedeagus and bulbus ejaculatorius, Mt Panié [NZAC prep. L 872]. Fig. 10. T. liparodes PT male, genitalia, Mt Panié [NZAC prep. L 880]. (Scales by Figs 4 & 5 repesent 1.0 mm; those by Figs 6, 8 – 10, 0.5 mm.)

FIGURES 11 – 15. Fig. 11. T. paniense, PT male, genitalia, Mt Panié [NZAC prep. L 878]. Fig. 12. T. nigrilinea, female genitalia, Baie des Tortues [NZAC prep. 866]. Fig. 13. T. liparodes, PT female genitalia, Mt Panié [NZAC prep. L 873, 874]. Fig. 14. T. paniense PT female genitalia: ductus and corpus bursae, Mt Panié [NZAC prep. 877]. Fig. 15. T. paniense PT female genitalia: ovipositor, sterigmal and limen structure [NZAC prep. L 877]. (All scales represent 0.5 mm.)