Oncophyes Marshall

Yunakov, Nikolai, 2012, 3535, Zootaxa 3535, pp. 1-104 : 45-46

Oncophyes Marshall

Marshall 1942: 20. Type species: Ellimenistes amoenus Hartmann, 1904 , by original designation.

Diagnosis. Similar to Cadoderus in structure of head and general appearance, but differs by large size: [BL: 4.02–5.69 (4.96) mm, BW: 1.80–2.80 (2.38) mm, BH: 1.62–2.60 (2.17) mm]; funicle entirely squamose; coloration consisting of red-brown background and striped pattern formed by brown, cupreous and white scales; elytral intervals 1, 2, 4, 6 with distinct row of setae; subgenae and area of occipital suture bare, glabrous; rostrum as long as wide or weakly elongate [RL/RW: 1.00–1.10 (1.05)]; eyes flat, delimited by glabrous convex ring; vertex narrow; funicular segment 1 shorter than 2nd; posterior setal fringe consists of spatulate setae, partly hidden by posterior edge of pronotum; subscutellar callosity very broad, glabrous, without tubercles, not hidden by scales; interior edge of protibiae and mesotibiae weakly C-shaped, with large teeth in distal portion; internal sac with spiculate fields and small aggonoporium consisting of two curved lobes connected proximally by transverse bridge and long, slender ventral process; female tergite 8 narrowlysubtrapezoid, apically acute.

Description.

Measurements. BL: 4.02–5.69 mm, BW: 1.8–2.8 mm, BH: 1.62–2.60 mm.

Vestiture. Body densely covered with overlapping, recumbent, round, flat scales, setose. Epifrons covered with erect scales. Setae of head and body slender, truncate, form rows along lateral edges of epifrons (suberect) and above the eyes (erect). Subocular row consists of 2–3 setae. Pronotum evenly setose, without anterior setal fringe. Posterior setal fringe consists of spatulate setae, partly hidden by posterior edge of pronotum. Elytral intervals 1, 2, 4, 6 (or 2, 4, 6) with distinct row of setae. Anterior half of elytra with rather short suberect setae (half as long as interval width), posterior declivity with rather long, strongly erect, thick, acute or truncate spatulate setae (as long as interval width). Ventral side of head capsule (except hls-ptp callosity), thorax and abdomen very densely squamose. Anterior part of pterygia, subgenae and area of occipital suture bare, glabrous. Antennal scape setose and densely squamose; scales round; setae long, slender, acute, and erect. Funicle setose and squamose; scales elongate; setae suberect, as long as funicular segment 7. Club densely tomentose. Femora and tibiae external and internal surfaces covered with overlapping scales and suberect, slender, acute setae. Male metatibiae with small grooming brush; hairs short. Tarsi setose and squamose.

Coloration. Integument of body brown to deep-brown; limbs deep-brown. Background red-brown; striped pattern formed by brown, cupreous, white or green scales. Dark and pale scales both with slightly pronounced pearl shine. Head dorsal surface cupreous-brown, without stripes, temples green or red-brown. Pronotum with cupreousbrown background scaling and 4 distinct longitudinal deep-brown stripes (2 thin median stripes and 2 broad lateral stripes). Elytra background scaling deep-brown or cupreous with white V-shaped transversal band or cruciform spot at the middle and large median cupreous spot; dark-with cupreous stripes at the elytral intervals 2 and 4 or interval 1 cupreous. Ventral side cupreous or whitish-cupreous with scattered greenish scales. Femora and tibiae brown-cupreous, femora with transverse brown band in the middle.

Head. Rostrum as long as wide or weakly elongate [RL/RW: 0.96–1.10], parallel-sided. Pterygia not extended beyond contour of rostrum. Antennal sockets dorsal. Antennifers entirely visible in dorsal view. Lateral carinae convex. Epifrons very narrow, parallel-sided, at the level of antennal articulation 1.78–2.00 x narrower than vertex, distinctly sloping at sides, with convex median carina (completely concealed by scales). Transverse sulcus deep, concealed by dense scaling at dorsal surface and at sides. Anterior portion of epifrons not steep, without median sulcus. Frons moderately declivous, glabrous, bare, with 4 frontal setae. Epistome convex, broad, arcuate, delimited by narrow carina, bears 2 tiny epistomal setae. Prementum with 2 setae. Mandibles with 3 setae. Eyes sublateral, large, orbicular, flat, delimited by glabrous convex ring [FW/ELD: 0.67–0.86]. Vertex narrow, flat. Frontal fovea elongate, partly concealed by scales.

Antennae. Scape extended behind anterior edge of pronotum, reaches anterior 1/3, weakly, evenly curved, thick. Funicular segment 1 shorter than 2nd; segments 3–7 oblong (3rd—1.50–1.54 x as long as wide; 7th –1.10–1.25 x as long as wide). Club broadly-spindle-shaped.

Thorax. Pronotum strongly transverse [PL/PW: 0.80–0.89], evenly slightly convex at sides, constricted at apex, widest at middle. Lateral longitudinal carina significantly developed to weak or absent. Disc weakly convex longitudinally and transversally. Tergosternal suture complete. Metanepisternal suture obsolete posteriorly.

Elytra. Cordate, anterior edge arcuate, vertical, narrowly rounded [EL/EW: 1.30–1.40]; disc in male moderately in female strongly convex [EL/BH: 1.40–1.58]; posterior declivity evenly sloping; elytral apex narrowly rounded, almost acute. Subscutellar callosity broad to oblong-eggshaped, weakly or strongly convex, without tubercles. Elytral intervals flat.

Legs. Femora unarmed, clavate, swollen in middle part. Protibiae and mesotibiae curved, not widened at the apex. Interior edge weakly C-shaped with large teeth in distal portion. Metatibiae with large teeth on interior edge. Mucro well developed, thorn-shaped, acute. Setal comb of protibiae consists of sparse 8–9 black short setae not extending beyond external edge of tibiae, partly hidden by spatulate setae. Bevel of metatibiae narrowly enclosed; its surface bare. Tarsi robust; setose pelma well developed; tarsomere 2 as long as wide; tarsomere 3 with two wide lobes; tarsomere 5 of metatarsus extending beyond apical lobes of 3rd by length of the last one. Claws connate in basal half.

Abdomen. 1st and 2nd ventrites fused but with distinct immovable suture. Posterior margin of 1st ventrite arcuate. 2nd ventrite 2 x as long as 3rd one, posterior margin of 2nd ventrite straight. 5th ventrite in male flat, in female weakly convex near the apex; apical edge in male broadly rounded, in female narrowly rounded.

Male genitalia. Aedeagus heavily sclerotized. Median lobe parallel-sided, tubular, in apical 1/3 evenly narrowed anteriorly; apex lanceolate, acute, strongly with strong median arcuate sharp carina; 0.7 x as long as apodeme. Internal sac with spiculate fields and small aggonoporium consisting of two curved lobes connected proximally by transverse bridge and long ventral slender process. Ostium lanceolate, strongly sclerotized. Ligulae membranous. Parameres long, fused in basal half, basal piece of tegmen narrow, tegminal apodeme 0.5 x as long as apodemes of median lobe.

Female genitalia. Coxites heavily sclerotized, dorso-ventrally curved, with heavily sclerotized dorsal baculi, evenly covered with pores, without sensilla. Styli well developed, stick-shaped, bearing 1 seta. Spermatheca moderately sclerotized, with short broad ramus, collum slender, as long as ramus; corpus large, strongly swollen; cornus slender, extended beyond corpus. Spermathecal duct antirely tubular or bubble-shaped at distal end ( Figs 353, 354). Tergite 8 narrowly-subtrapezoid, apically acute, with curved fringe of multiple tenuous, short setae at the apex. Sternite 8 thick. Lamella sharply narrowed and acute, without setae, knife-shaped, heavily sclerotized. Apodeme thick, caput large.