Landouria moussoniana ( Martens, 1867 ) Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 53-54

Landouria moussoniana ( Martens, 1867) comb. nov.

Figs 55 , 75 ; Tables 3–4

Helix sumatrana var. moussoniana Martens, 1867: 266 (“Oestliches Java, im Tengergebirge bei Wonosari”).

Landouria rotatoria – van Benthem Jutting 1950: 461 (in part). (not Pfeiffer, 1842)

Diagnosis

Landouria moussoniana is characterized by a tightly coiled, keeled shell with a hardly eccentric, very wide umbilicus.

Material examined

Syntypes

INDONESIA • 6 spec.; East Java, Wonosari ; 7°54′ S, 112°49′ E; ZMB 109902 .

Description

SHELL ( Fig. 75 ; Tables 3–4 ). Depressed conical, with 5–6.25 slightly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes not observed; tuberculate around umbilicus; brownishcorneous; body whorl keeled; aperture oval; upper insertion of peristome hardly or distinctly descending; peristome expanded, reflexed and slightly thickened; umbilicus very wide, comprising 37–38% of shell diameter, hardly obscured by columellar edge.

GENITALIA. Unknown.

Remarks

Martens (1867) recorded L. moussoniana from Wonosari in the Tengger Mountains together with L. smimensis and another form of Landouria that he included in his L. intumescens . Landouria moussoniana differs from L. intumescens in having a more tightly coiled, less sharply keeled shell with a wider umbilicus (U/D 0.37–0.38 vs 0.22–0.34 in L. intumescens ).

Distribution

Landouria moussoniana is endemic to the Tengger Mountains in East Java (Fig. 55).

Figures

Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867).

Figs 72–75. Shells of species of Landouria from Java. 72. L. intumescens (Martens, 1867), East Java, Surabaya (lectotype, ZMB 5160a). 73. L. intumescens, East Java, Guwoterus (MZB 19207). 74. L. intumescens?, East Java, Wonosari (paralectotype, ZMB 230704). 75. L. moussoniana (Martens, 1867), East Java, Wonosari (syntype, ZMB 109902). Scale bar: 5 mm.