Hemyda Robineau-Desvoidy, 1830

Wang, Qiang, Zhang, Chuntian & Wang, Xinhua, 2015, Review of the Hemyda Robineau-Desvoidy of China (Diptera: Tachinidae), Zootaxa 4040 (2), pp. 129-148 : 130-131

1. Hemyda Robineau-Desvoidy, 1830

Hemyda Robineau-Desvoidy, 1830: 226 . Type species: Hemyda aurata Robineau-Desvoidy, 1830 , by monotypy. Type locality: USA, Pennsylvania, Philadelphia.

Main references: Herting & Dely-Draskovits 1993: 432 (Palearctic catalogue); Ziegler & Shima 1996: 441 (Russian Ussuri Tachinidae ); Tschorsnig & Richter 1998: 752 (key to Palearctic genera); O’Hara & Wood 2004: 219 (Nearctic catalogue); O’Hara et al. 2009: 128 (Chinese catalogue); Cerretti et al. 2012: 8 (MOSCHweb interactive key).

Description. Male and female. Body length: 7.0−11.0 mm.

Head: Eye bare; parafacial bare; face flat, wider than eye width in frontal view. Lower facial margin flat or at most weakly protruded forward; frontal vitta broad, widened posteriorly; ocellar triangle large; gena at most 1/10 of eye height, genal dilation slender, with long black setulae; occiput slightly convex; hind margin of eye usually indented. Inner vertical seta strong or fine, outer vertical seta fine or slightly longer than postocular setae; proclinate and reclinate orbital setae absent in male, 0−2 proclinate and 0−2 reclinate orbital setae in female; ocellar setae strong or fine. Occiput with several short black setulae and dense pale setulae; 2−4 fine postocellar setae; postocular setae absent on lower 1/3−3/5 of eye hind margin. First flagellomere 2.7−3.5 times as long as pedicel; arista almost bare; 2nd aristomere 1.0−2.2 times as long as wide. Prementum short, at most twice as long as its diameter; palpus slender, weakly swollen near apex.

Thorax: Prosternum and proepisternum bare; postpronotal lobe with 3 setae arranged in a straight line; 0−2 presutural and 1−2 postsutural acrostichal setae; 3 presutural and 3 postsutural dorsocentral setae; 0 presutural and 2 postsutural intra-alar setae, anterior postsutural seta at same level as anteriormost postsutural dorsocentral seta or slightly anterior, and posterior postsutural seta at same level or slightly posterior to level of second postsutural dorsocentral seta; 2 supra-alar setae; 1−2 katepisternal setae; anepimeral seta hair-like; katepimeron bare; scutellum with 3 pairs of strong marginal setae, apical ones crossed. Wing hyaline, evenly tinged with pale brown; costa setulose dorsally, and ventrally first, second and apex of third costal sectors setulose; costal spine not differentiated; wing veins bare except basal node of vein R4+5 with 3−6 fine setulae dorsally and ventrally; cell r4+5 open. Legs black to dark brown or partly yellow; fore tibia with 2 preapical dorsal setae, anterodorsal seta absent, and 0−2 posterior setae; mid tibia with 0−2 ventral setae; hind tibia with 2 apical dorsal setae, 1 apical anteroventral seta, and without an apical posteroventral seta; male claws and pulvilli long, subequal or slightly shorter than 4th and 5th tarsomeres combined, shorter than 5th tarsomere in female.

Abdomen ovate or subcylindrical, broadly yellow, without median discal setae and with or without pruinosity; syntergite 1+2 excavated on at most basal 1/2 and with 2 median marginal setae; tergite 5 strongly curved anteroventrally; sternites well exposed. Male terminalia: Sternite 5 broad, with posteromedian cleft usually Ushaped or V-shaped, broadly or weakly excavated posteromedially, without a membranous area anterior to median cleft, usually with fine setulae posteriorly; syntergite 6+7 shorter than 5th tergite; epandrium usually small and narrow in caudal view, with fine setulae, anterior margin straight, basal median portion narrow, lateral margin usually short in lateral view; cerci short in caudal view, usually with fine short setulae on inner lateral margin, separated on more than upper half dorsally; surstylus long and greatly enlarged posteriorly in lateral view, with fine short setulae laterally and posteriorly; hypandrium broad in lateral view, hypandrial arm long, phallapodeme long, with phallic guide (intermedium); pregonite and postgonite fused, broad and long in lateral view; epiphallus and phallus elongate, apex pointed; ejaculatory apodeme short. Female terminalia: Syntergite 6+7 longer than tergite 5, usually with long and erect setulae, ventral portion of tergite 6 with a long finger-like protrusion, bent posteriorly; tergite 7 divided into two broad lobes laterally, bent anteriorly; sternite 7 short; tergite 8 narrow, posteriorly divided into two elongate lobes, with fine setulae of variable length, rounded or pointed at apex; sternite 8 long, forming a piercer.

Remarks. Hemyda is close to Phania by sharing the following external character states: Eye, parafacial and arista bare, scutellum with 3 pairs of marginal setae, wing cell r4+5 open or at most with a short petiole, hind tibia without a preapical posteroventral seta, postmetacoxal area sclerotized, middorsal depression of abdominal syntergite 1+2 not reaching posterior margin, sternites well exposed, and terminalia of male and female prominent and well visible in lateral view. Hemyda differs from Phania by having pale setulae on the occiput, at most with several black setae on upper part behind postocular setae ( Phania with black setulae only), lower facial margin usually not visible in lateral view (always visible in lateral view in Phania ), hind margin of eye indented, prementum short, at most three times longer than broad (in Phania at least five times).