Zeadalmodes Nomura and Leschen, 2015

Nomura, Shûhei & Leschen, Richard A. B., 2015, Myrmecophilous Pselaphinae (Coleoptera: Staphylinidae) from New Zealand, The Coleopterists Bulletin 69 (1), pp. 121-152 : 121-152

Zeadalmodes Nomura and Leschen , new genus ( Figs. 12–17 )

Type species: Zeadalmodes myrmecophilus Nomura and Leschen , new species

Diagnosis. Body large (2.72–3.18 mm), robust and elongate. Head narrower than prothorax and nearly circular in dorsal view, frons weakly concave, vertex slightly convex, with pair of welldemarcated, setose, round dorsal tentorial pits;

138 THE COLEOPTERISTS BULLETIN 69(1), 2015

posterior constriction not notched and lacking lateral fovea; postgenae large and rounded, venter bearded ( Fig. 13B ). Antennae short and thick, weakly thickened towards apex, with insertions hidden in dorsal view; antennal tubercles weakly defined. Maxillary palpi short and thick, palpomere IV largest, nearly fusiform.

Pronotum about as long as head, gently convex and nearly circular, with pair of well-developed basilateral foveae, 3 longitudinal sulci (pair of lateral longitudinal sulci and median longitudinal sulcus), and well-demarcated bisinuate antebasal sulcus ( Fig. 13A ). Elytra slightly wider than long, weakly convex, rounded on humeri; each elytron with 2 basal foveae, humeral denticle, short longitudinal sulcus extending from outer basal fovea, and adsutural sulcus extending from inner basal fovea ( Fig. 13F ). Prosternum without lateral or median longitudinal carinae. Ventral foveae simple, those present as follows: gula (head); lateral procoxal (prosternum); small prepectal, lateral mesoventral, fused median mesoventral and lateral mesocoxal (mesoventrite); lateral metaventral (metaventrite); weak mediobasal and basolateral (abdominal sternite III). Legs short and stout; 1 tarsal claw present.

Abdomen elongate and subparallel-sided, with tergites equal in length, sternite III very short, and large, V-shaped projection between metacoxae; IV longer than III, large, broadened posteriorly, with pair of narrow paratergites on lateral sides; V about as large as IV, broadened posteriorly at sides.

Remarks. Zeadalmodes is easily separated from other trichonychine genera by having a large and robust body, a nearly circular pronotum with a welldemarcated bisinuate transverse sulcus, and a subparallel-sided abdomen.

Etymology. The new generic name is formed from the prefix “ Zea- ” meaning “of New Zealand ” and combined with the generic name “ Dalmodes ”.

Figures

Fig. 12. Zeadalmodes species, habitus. A) Z. myrmecophilus, male holotype, B) Z. brevis, female paratype. Scale = 1 mm.

Fig. 13. Zeadalmodes myrmecophilus, male. A) Head and prothorax, dorsal view, B) Head and prothorax, in ventral view, C) Right maxillary palpus, ventral view, D) Right antenna, ventral view, E) Meso- and metathoraces, ventral view, F) Left elytron, dorsal view, G) Abdomen, dorsal view, H) Abdomen, ventral view.

Fig. 14. Zeadalmodes myrmecophilus, male. A) Tergite VIII, B) Sternite VIII, C) Tergite IX (two lateral plates) and sternite IX (middle plate), D) Genitalia, ventral view, E) Genitalia, lateral view. Scale = 0.1 mm.

Fig. 15. Zeadalmodes myrmecophilus, female. A) Tergite VIII, B) Sternite VIII, C) Genital plate in ventral view. Scale = 0.1 mm.