Teloganodes, Eaton, 1882

Kluge, Nikita, 2023, Reclassification of Teloganodes Eaton 1882 (including subgenera Dudgeodes Sartori 2008 and Derlethina Sartori 2008) with a new species from Sulawesi (Ephemeroptera, Teloganodidae), Zootaxa 5244 (6), pp. 527-552 : 529-530

Autapomorphies of Teloganodes

(1) On tergalii beginning from the pair III (either pairs III–V, or pairs III–IV, or pair III only), dorsal lobe is deeply cleft ( Figs 1 , 51–53, 55–56 ) (unique autapomorphy). Other, non-unique features of tergalii structure are: tergalius I is absent; tergalius II is operculate, i.e. with dorsal lobe convex, larger than others and never cleft; tergalii II and next ones (either II–V, or II–IV, or II–III) retain the two-branched ventral lobe bearing processes on both its branches (that is initial for Ephemerelloidea ) ( Figs 1 , 50–57 ).

(2) Larval, subimaginal and imaginal paracercus is vestigial, 1-segmented. The same in some other mayflies; among Ephemerella/fg1, the same in Dicercomyzon Demoulin 1954 only.

(3) In subimago, paired lateral pigmented area of mesonotum (termed here latero-posterior scutal chromozone) occupies the whole sublateroscutum, the whole submedioscutum and a latero-posterior part of medioscutum, leaving a non-pigmented median area (termed here achromozone) in a form of unpaired median stripe narrowing posteriorly; the latero-posterior scutal chromozone is separated from the achromozone by a distinct, evenly arched line termed here latero-posterior chromozonal suture ( Figs 27–28 ). The area anteriad of the achromozone is termed here anterior scutal chromozone. Color of the anterior scutal chromozone, the lateroposterior scutal chromozones and the achromozone vary among species, but shape of their borders is constant for Teloganodes . Microtrichia cover most part of scutum, being absent on lateroscutum and posterior scutal protuberances ( Fig. 99 ). In contrast to Ephemerellidae and the most primitive representatives of Pantricorythi, the lateroparapsidal sutures of subimago and imago of Teloganodes are not curved laterally, but converge with the medioparapsidal sutures ( Kluge 2004: p. 294, character 10 of Ephemerella/fg1) ( Fig. 28 ).

Figures

FIGURE 1. Tergalii figured by Kluge (2004: figs 96E–H) and two classifications of Teloganodes. E, tergalii of T. (Teloganodes) kodai; F, tergalii of T. (Dudgeodes) romani (= Teloganodes/fg1 sp. T4: Kluge 2004); G, tergalii of T. (Dudgeodes) jacobusi (= Teloganodes/fg1 sp. T2: Kluge 2004); H, tergalii of T. (Derlethina) eloisae (= Teloganodes/fg1 sp. T3: Kluge 2004).

FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype).

FIGURES 27–30. Teloganodes (Dudgeodes) celebensis. 27–28, left half of subimaginal exuviae of mesonotum; 29–30, hind wing in falling and transmitted light. Abbreviations: acz, achromozone; cz.a, anterior chromozone; cz.l-p, latero-posterior chromozone; cz.s, chromozonal suture; LPs, lateroparapsidal suture; MPs, medioparapsidal suture; MS, medioscutum; SMS, submedioscutum.

FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light.