Coenosia Meigen

Xue, Wan-Qi & Zhu, Yu, 2006, Three new species of Coenosia Meigen (Diptera: Muscidae) from P. R. China, with a key to the Chinese species of the genus, Zootaxa 1326, pp. 1-16 : 2-8

Key to species of the genus Coenosia Meigen from China (males)

1. Dc 2+3 ........................................................................................................................... 2

­ Dc 1+3 ........................................................................................................................... 6

2. Femora black, at most apex and base yellow ................................................................ 3

­ Mid and hind femora yellow or mostly yellow ............................................................. 5

3. Thorax with grayish white pruinosity, without vittae; hind tibia without pd. ................. ............................................................................................ C. lacteipennis (Zetterstedt)

­ Thorax with brown pruinosity, with fuscous longitudinal vittae; hind tibia with 1–2 small pd ......................................................................................................................... 4

4. Parafacial about three­fifths width of antenna; abdominal tergites 3–5 with spots, cerci asymmetrical in posterior view ............................. C. ansymmectrocerca Xue and Feng

­ Parafacial about one­fourth width of antenna; abdominal tergites 2–4 with spots, cerci symmetrical in posterior view ................................. C. brunneipennis Cui, Xue and Liu

5. Mid and hind femora entirely yellow; pedicel and flagellomere 1 reddish brown at base .......................................................................................................... C. scissura Ma

­ Mid and hind femora gray near apex; antenna black ................................. C. pauli Pont

6. Costa ending slightly beyond vein R4+5 ......................................................................... 7

­ Costa ending in vein M ................................................................................................. 8

7. 1 proepisternal seta; mid femur and tibia yellow, hind femur and tibia blackish brown .................................................................................................... C. cornectens (Hennig)

­ 2 proepisternal setae; mid and hind legs black ............................ C. oligochaeta Hennig

8. Hind tibia with apposed av and ad ................................................................................ 9

­ Hind tibia without apposed av and ad ......................................................................... 16

9. Antenna with at least apex of flagellomere 1 yellow.. ............................................... 10

­ Antenna entirely black or fuscous............................................................................... 12

10. Antenna entirely yellow, frontal vitta with bright silvery pruinosity. C. attenuata Stein

­ At least most of pedicel brown, frontal vitta without silvery pruinosity..................... 11

11. Femora with dense fine ventral hairs, especially on mid and hind legs.......................... .............................................................................................................. C. strigipes Stein

­ Femora without fine hairs on ventral surface .... C. strigipes bannaensis Xue and Tong

12. Legs mostly yellow, femora brown on dorsal surface ............................. C. exigua Stein

­ At least fore femur black............................................................................................. 13

13. At least mid and hind femora yellow on basal one­fourth .......................................... 14

­ Mid and hind femora mostly black or fuscous............................................................ 15

14. Mid and hind femora yellow on basal two­thirds; frontal vitta distinctly with gray pru­ inosity ............................................................................. C. breviaedeagus Wu and Xue

­ Mid and hind femora yellow on basal one­fourth; frontal vitta black, without pruinos­ ity ...................................................................................... C. albisquama Xue and Tong

15. Frontal vitta with bright gray pruinosity; thorax and abdomen with grayish pruinosity ........................................................................................................... C. humilis Meigen

­ Frontal vitta with dense dark brown pruinosity; thorax and abdomen with fuscous pru­ inosity .................................................................. .. C. attenuata brunnea Xue and Tong

16. First and second tarsomeres of mid and hind legs distinctly broadened, pulvilli of mid and hind legs longer than those of fore leg ..................................... C. dilatitarsis Stein

­ Characters not as above...............................................................................................17

17. Hind tibia with 2 coarse posterior median spines; body black, abdominal spots indis­ tinct.............................................................................................. C. chinensis (Malloch)

­ Hind tibia without coarse spines .................................................................................18

18 Hind tibia with 2 preapical d ....................................................... C. intermedia (Fallén)

­ Hind tibia with only 1 preapical d ...............................................................................19

19. Lower calypter extremely small, shorter than half length of the upper one................20

­ Lower calypter similar to or longer than the upper one. .............................................31

20. Femora mostly yellow, only the dorsal surface of fore femur and the distal one­fourth of mid and hind femora fuscous............................................... .. C. orbimacula sp. nov.

­ Femora mostly black or fuscous..................................................................................21

21. Hind tibia with 2 pd (1 fine)........................................................................................22

­ Hind tibia only with 1 pd .............................................................................................28

22. Tibiae entirely yellow............................................................. ... C. fengi Yang and Zhao

­ Tibiae fuscous or black, except for base yellow.......................................................... 23

23. Femora with many fine ventral hairs on basal half; genal height at least eighth of eye height. ................................................................................... C. fimbripeda Wu and Xue

­ Femora without fine ventral hairs on basal half; genal height at least one­sixth of eye height...........................................................................................................................24

24. Abdomen with a median rectangular spot............. C. mimolongipeda Li, Feng and Xue

­ Abdomen with triangular spots laterally or without spots .........................................25

25. Genal height about one­sixth of eye height................................................................. 26

­ Genal height about one­fourth of eye height...............................................................27

26. Frons about one­third width of head, parafacial with dark brown pruinosity, flagellom­ ere 1 about 2.5 times as long as broad; palpus yellow; cerci and surstyli curved in S­ shape in profile .............................................................. C. obscuripennis Xue and Feng

­ Frons about two­fifths width of head, parafacial with gray pruinosity, flagellomere 1 about 1.5 times as long as broad; palpus black; cerci and surstyli very straight in pro­ file ................................................................. C. angulipunctata Xue, Wang and Zhang

27. Thorax with five gray vittae; hind tibia with 1 av, 1 ad ....... .. C. spinifemorata sp. nov.

­ Thorax with three gray vittae; hind tibia with 2 av, 2 ad ... ungulipardus Xue and Feng

28. Tarsi distinctly shorter than tibiae; sternite 5 with long dense hairs ............................... .................................................................................... C. brevimana Cui and Xue, 2001

­ Tarsi distinctly longer than tibiae; sternite 5 without long hairs ................................ 29

29. Parafacial about one­third width of flagellomere 1, palpus yellow on basal half; fore tibia yellow, mid and hind tibiae fuscous .............................. C. longipeda Wu and Xue

­ Parafacial about half width of flagellomere 1, palpus dark brown; tibiae entirely yel­ low or brownish yellow............................................................................................... 30

30. Genal height about one­third of eye height; hind tibia with 1 pd ... C. changjianga Xue

­ Genal height about one­fourth of eye height; hind tibia with 2 pd. .. C. shennonga Xue

31. Lower and upper calypters subequal in length............................................................ 32

­ Lower calypter longer, distinctly protruding beyond upper one ...............................40

32. At least most of mid and hind femora yellow. ............................................................33

­ Mid and hind femora entirely black or slightly yellow at apex. ................................. 38

33. Femora entirely yellow................................................................................................34

­ Mid and hind femora dark brown or black on distal one­fourth to one­third ............. 36

34. Body hairs mostly yellow; flagellomere 1 yellow on basal half, fuscous on distal half, about 2.7 times as long as broad, palpus yellow. ............ ... C. subflaviseta Xue and Cui

­ Body hairs mostly black; other characters varied. ......................................................35

35. Flagellomere 1 entirely yellow. ...................................................... C. mollicula (Fallén)

­ Antenna black, flagellomere 1 about twice as long as pedicel........................................ ............................................................................................. C. yulongxueshanensis Xue

36. Antenna brownish, parafacial narrower than half width of flagellomere 1.. ................... ................................................................................. C. unpunctata Xue, Yang and Feng

­ Antenna and palpus black; parafacial varied in width. ............................................... 37

37. Parafacial and flagellomere 1 subequal in width; fore femur black at apex, mid and hind femora black on distal one­third to half.................... C. octopunctata (Zetterstedt)

­ Parafacial about two­thirds width of flagellomere 1; fore femur mostly yellow except for the fuscous dorsal surface, mid and hind femora dark brown on distal one­fourth.. ..................................................................................................... C. carinata Cui and Li

38. Body length at least 4.5 mm ... ........................................................... C. hirsutiloba Ma

­ Body length shorter than 4.0 mm. ............................................................................... 39

39. Hind tibia without av; abdomen subequal in length to thorax ........................................ ............................................................................................ C. nigrimixta Feng and Xue

­ Hind tibia with av; abdomen longer than thorax................. C. subgracilis Xue and Cui

40. Flagellomere 1 with cone­like process at apex. ..........................................................41

­ Flagellomere 1 without cone­like process at apex ...................................................... 48

41. 1 proepisternal seta...................................................................................................... 42

­ 2 proepisternal setae.................................................................................................... 45

42. Coxae and femora entirely black or fuscous ...............................................................43

­ Coxae and femora colour not as above ....................................................................... 44

43. Mid tibia with ad; legs entirely black..................................... C. flavimana (Zetterstedt)

­ Mid tibia without ad; apex of femora yellow.......................... C. pulicaria (Zetterstedt)

44. Frontal vitta distinctly with yellowish white pruinosity; mid and hind coxae brown; abdomen with one spot on each tergite.. ........................................ C. ambulans Meigen

­ Frontal vitta without pruinosity; mid and hind coxae brownish; abdomen yellow on basal half, without spots ............................................................... C. flaviambulans Xue

45. Antenna, face and gena all yellowish white; legs entirely yellow; body hairs yellowish ...................................................................................................... C. aurea Xue and Cui

­ Antenna, legs and body hairs at least partly black ...................................................... 46

46. Antenna yellow, parafacial about half width of flagellomere 1; legs mostly yellow...... ............................................................................................................... C. erdosica Tian

­ Antenna black or fuscous; parafacial varied; legs black .............................................47

47. Parafacial about three­fourths width of flagellomere 1; the latter about 1.6 times as long as pedicel... ........................................................................... C. parva Xue and Cui

­ Parafacial about one­third width of flagellomere 1; the latter about 2.3 times as long as pedicel. .............................................................................................. C. grisella Hennig

48. 1 proepisternal seta...................................................................................................... 49

­ 2 proepisternal setae. .................................................................................................. 52

49. Mid tibia with ad ... ...................................................................................................... 50

­ Mid tibia without ad .................................................................................................... 51

50. Flagellomere 1 yellow on basal half; face, parafacial and gena all grayish white.......... ....................................................................................................... ... C. incisurata Wulp

­ Flagellomere 1 entirely fuscous; face, parafacial and gena all yellowish. C. zhaoi Xue

51. Flagellomere 1 black, about 3 times as long as pedicel .................................................. .................................................................................... C. testacea (Robineau­Desvoidy)

­ Flagellomere 1 yellow, about twice as long as pedicel ................ C. xanthocera Hennig

52. Flagellomere 1 entirely yellow. ................................................................................... 53

­ Antenna black or only flagellomere 1 yellow on basal half; trochanters yellow at least ..................................................................................................................................... 54

53. Flagellomere 1 acute­angled at apex, parafacial about three­fourths width of flagellomere 1 ............................................................................... C. flavicornis (Fallén)

­ Flagellomere 1 obtuse­angled at apex, parafacial about two­fifths width of flagellom­ ere 1 ............................................................................................. C. subflavicornis Hsue

54. Femora black or fuscous except for apex ................................................................... 55

­ Mid and hind femora yellow at least on basal half ..................................................... 57

55. Tibiae black except for basal part ......................................... C. nigricornis Wu and Xue

­ Tibiae entirely yellow..................................................................................................56

56. Mid femur with av, mid tibia with 1 ad, hind tibia with 1–2 pd. .................................... .................................................................................................... C. amplicauda sp. nov.

­ Mid femur without av, mid tibia without ad, hind tibia without pd .. .............................. .................................................................................... C. 1ongiquadrata Xue and Wang

57. Fore femur black at least on basal two­thirds ............................................................58

­ Fore femur mostly yellow or distal two­thirds yellow on dorsal surface.................... 67

58. Fore femur fuscous on basal two­thirds, mid and hind femora entirely yellow.............. ............................................................................................. C. monticola Xue and Zhao

­ Fore femur black except for apex, mid and hind femora dark brown on distal one­ fourth. .......................................................................................................................... 59

59. Mid tibia without ad .................................................................................................... 60

­ Mid tibia with ad ......................................................................................................... 63

60. Mid femur yellow on basal one­fifth to one­fourth, hind tibia without av .................. 61

­ Mid femur yellow on basal two­thirds at least, hind tibia with av .. ............................ 62

61. Fore tarsus yellow, abdomen distinctly with middle vittae and lateral spots.................. ............................................................................................. C. flavimixta Feng and Xue

­ Tarsi black, abdomen without middle vittae ........................... C. grandis Xue and Zhao

62. Frontal vitta orange, palpus brownish, parafacial about half width of flagellomere 1; scutum without spots ........................................... C. unguligentilis Xue, Yang and Feng

­ Frontal vitta black, palpus black, parafacial about one­third width of flagellomere 1; scutum with 2 brown spots. ........................................................... C. albicornis Meigen

63. Hind tibia with 1 developed pd, about two­thirds length of the prst dc .......................... ............................................................................................................ C. xuei Cui and Li

­ Hind tibia without pd ................................................................................................... 64

64. Tarsi yellow, mid and hind femora brownish in distal one­fourth on dorsal surface; body length 4.0 mm ............................................................ C. fanjingensis Cui and Xue

­ Hind tarsus fuscous or black; body length varied. ...................................................... 65

65. Legs yellow except for coxae and tarsi black ....... C. campestris (Robineau­Desvoidy)

­ Leg colour not as above .............................................................................................. 66

66. First tarsomere of fore leg yellow, first tarsomere of other legs dark brown or black, mid and hind femora black on basal half. ........................................... C. verralli Collin

­ First to fourth tarsomeres of fore leg and first tarsomere of mid leg yellow, mid femur yellowish brown on distal one­fourth, hind femur fuscous on distal one­third .............. ................................................................................................. C. pygmaea (Zetterstedt)

67. Fore femur fuscous on distal two­thirds of dorsal surface, mid and hind femora fus­ cous on distal one­fourth .................................................................. C. striolata Hennig

­ Femora entirely yellow or fore femur slightly brownish on dorsal surface, mid and hind femora brownish at apex on dorsal surface......................................................... 68

68. Apical scutellar seta distinctly shorter than the basal one........................................... 69

­ Apical scutellar seta longer than the basal one. .......................................................... 70

69. Wing narrow on basal half; lower facial margin and the anterior margin of frons in the same vertical plane ............................................................................ C. pedella (Fallén)

­ Wing not narrow on basal half; lower facial margin behind the anterior margin of frons in profile .................................................................................. C. griseiventris Ringdahl

70. Basal scutellar seta less developed than the apical one, about three­fourths length of the latter .................................................................................... C. nigrifrons Cui and Li

­ Basal scutellar seta as developed as the apical one..................................................... 71

71. Hind tibia with pd ........................................................................................................ 72

­ Hind tibia without pd ................................................................................................... 75

72. Hind tibia with 1 developed ad; 1 fine hair before prst dc, about one­third length of the latter. ............................................................................................... C. luteipes Ringdahl

­ Hind tibia with 2 ad; other character varied................................................................ 73

73. Femora entirely yellow; thorax with indistinct vittae... .................. C. lacustris Schnabl

­ Hind femur with dark brown spots on dorsal surface; thorax distinctly with two dark vittae. ........................................................................................................................... 74

74. Abdominal tergites 7–9 subequal in length to tergite 5; tarsi fuscous ............................ .................................................................................... C. punctifemorata Cui and Wang

­ Abdominal tergites 7–9 shorter than tergite 5; tarsi brownish yellow ............................ ........................................................................................ C. taibaishanna Cui and Wang

75. Mid tibia with ad ... ...................................................................................................... 76

­ Mid tibia without ad .................................................................................................... 77

76. Abdomen cone­shaped, tergites 7–9 distinctly longer than tergite 5 .............................. ........................................................................................................ C. conica Cui and Li

­ Abdomen normal in shape, tergites 7–9 shorter than half length of tergite 5 ................. ........................................................................................................... C. paludis Tiensuu

77. Flagellomere 1 yellow on basal half; hind femur fuscous on distal one­fourth of dorsal

surface. ............................................................................................... C. pudorsa Collin ­ Antenna entirely black; hind femur entirely yellow.................................................... 78

78. Mid and hind femora with developed pv on distal one­third.. ............ C. sallae Tiensuu

­ Mid and hind femora without pv on distal one­third................................................... 79

79. Frontal triangle reaching the anterior margin of frons, palpus brownish yellow on basal half, fuscous on distal half... ................................................. C. mandschuricda Hennig

­ Frontal triangle reaching one­third to half way to frons, palpus not as above............80

80. Frontal triangle reaching one­third way to frons, palpus entirely fuscous; femora with­ out spots... .................................................................... C. spatuliforceps Xue and Zhao

­ Frontal triangle reaching half way to frons, palpus black; femora with annular black spots at apex .................................................................... C. lagenicauda Xue and Zhao