Angustopila Jochum

Páll-Gergely, Barna, Jochum, Adrienne & Asami, Takahiro, 2017, Three New Species And A New Genus Of Hypselostomatidae (Gastropoda: Pulmonata) From Cong Troi Cave, Northern Vietnam, Acta Zoologica Academiae Scientiarum Hungaricae 63 (3), pp. 327-341 : 329-333

Genus Angustopila Jochum , SlapniketPáll- Gergely, 2014

Angustopila Jochum , SlapniketPáll- Gergely, In: JOCHUM etal., ZooKeys 410: 26.

Typespecies: Systenostomatamlod PanhaetBurch 1999, byoriginaldes- ignation.

Content: concava (ThompsonetUpatham, 1997) , dominikae Páll- Gergely etHunyadi, 2015, elevata (ThompsonetUpatham, 1997) , huoyani Jochum , SlapniketPáll- Gergely, 2014, fabella Páll- GergelyetHunyadi, 2015, singuladentis InkhavilayetPanha, 2016 , subelevata Páll- GergelyetHunyadi, 2015, stochi Páll- GergelyetJochum, sp. n., szekeresi Páll- GergelyetHunyadi, 2015, tamlod (PanhaetBurch, 1999) .

Distribution: ThisgenusisdistributedinnorthernThailand, southern ChinaandnorthernVietnam ( Fig. 7).

Angustopilastochi Páll- GergelyetJochumsp. n.

( Figs 1C, 3A–H)

Typematerial: C03-N39 (localitycode), NorthernVietnam, HaGiangProvince, Quan

BaDistrict, TamSon, CongTroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, atthe entrance, goursofdrippingwater, leg. G. TOMASIN, 28.01.2010., HolotypeMZUFGC/49405; C04-N74 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, Tam Son, CongTroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, internalcanyon, goursof drippingwater, leg. G. TOMASIN, 29.01.2010., MZUFGC/49412 (figuredparatype, juvenile). (umbilicalside), G = sculptureofthebodywhorl.

Diagnosis: atiny, convexlyconicspecieswithanovate- subquadrateap- erture, atuberculatedparietalandalowpalataldenticle.

(umbilicalside), H = sculptureofthebodywhorl

Description: Shellminute, greyishwhite, veryfragile, spireconvexlyconicwithob- tuseapex; whorlsshouldered; spiretiltedslightlyleft; fineequidistantrowsofstriaeradiate fromthenuclearwhorl, startingatazoneofgranulosewrinklesandcontinuingontothe teleoconch; teleoconchsculpturefinelyretiformwithirregularly- spacedradialgrowthlines crossedbyfinerowsofspiralthreads; the 4.75 rounded, evenlyincreasingwhorlsaresepa- ratedbyadeepsuture; apertureovate- subquadrate; parietaldenticledeeply- set, consisting ofthreetuberclesrunninginside, thefirstconspicuouslylowerthanthesubsequenttwo; oppositeoftheparietaldenticleisasmall, deeply- set, lowpalatalplica; distalportionofbody whorlformsashortdetachedtuba; peristomethinatuppermostrightregion, basallyslightlyreflected; umbilicalzonehighlyreticulate; umbilicusnarrow; penultimatewhorlbulges decidedlybeyondtheplaneoftheaperturebyca, 1/6 themax. breadthofshell (lateralview).

Measurements (in mm): SH = 1.26; SW = 1.06; AH = 0.43; AW = 0.42 (holotype).

Differentialdiagnosis: Angustopilastochi sp. n. differsfromthesympa- tric A. subelevata by the larger shell and the presence of two denticles in the aperture. Three Angustopila speciespossesstwoaperturalteethsimilarto A. stochi sp. n., namely, A. huoyani , A. tamlod and A. dominikae . These species are considerably smaller than A. stochi . Moreover, A. huoyani , A. tamlod and A. dominikae bearamorepronounced, tuberculatedparietaldenticlecausingthe aperturetoappearheart-shaped. A. dominikae hasamoreaccentuatedpalatal denticleandasubglobose- conicalshell.

Etymology: ThisnewspeciesisdedicatedtoDr. FabioStoch, whosentthesamplesto

Dr. MarcoBodonwhokindlyprovidedusthematerial.

Distribution: Angustopilastochi sp. n. isknownfromsedimentsamples fromtheCongTroiCaveinNorthernVietnam.

Remarks: Thelastquarterwhorloftheholotype (theonlyknownadult shell) got damaged during the SEM imaging. Hence, these two shell frag- mentsaredepositedintheMZUF. Thisbreakageisprobablyduetoshellde- cayonaccountofitspreviouspreservationinformaldehyde (MarcoBodon, pers. comm.).

Angustopilasubelevata Páll- GergelyetHunyadi, 2015

( Figs 1B, 2A–G, 7E–F.)

Angustopila subelevata Páll- Gergely et Hunyadi, In: PÁLL-GERGELY et al. 2015, ZooKeys 523: 39–42, Fig. 4.

Materialexamined: C05-N5 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, TamSon, CongTroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, attheentrance, lakewithdrippingwater, leg. G. TOMASIN, 29.01.2010., 2 stronglycorroded shells, MZUFGC/49398; C03-N39 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, TamSon, CongTroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, tureheightandmultipliedby 100.

at the entrance, gours of dripping water, leg. G. TOMASIN, 28.01.2010., MZUF GC/49404 (1 shell); C04-N74 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, Tam Son, CongTroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, internalcanyon, goursof dripping water, leg. G. TOMASIN, 29.01.2010., Specimen 1 (figured shell), Specimen 2 (figured shell), additional 26 shells, MZUF GC/49411; Same data, HNHM 99733 / 1; Same data, PGB/2 .

Measurements (in mm): SH = 0.82–0.94; SW = 0.78–0.86; AH = 0.32–0.36; AW = 0.33–

0.35 (n = 12) ( Table 1).

Distribution: Angustopilasubelevata wasdescribedfromGuangxi, south- ernChina. Thelocalityreportedinthispaperrepresentsthefirstlocalityfrom Vietnam.

Remarks: TheVietnamesespecimensareslightlylargerthanthetype materialfromGuangxi, China (Min. SH: 0.83, maxSH: 0.91, mean: 0.87: Páll- Gergely etal. (2015), butnootherconchologicaldifferenceswerefound. The parietalcallusofoneoftheshellsformsatooth- likeprojection ( Fig. 7F).

Thematerialexaminedherehasspiralthreadsontheumbilicusandat the base of the shell in sync with PÁLL-GERGELY et al. (2015). PANHA and BURCH (2005) didnotobservethischaracteristicintheirinvestigationsofthemost similarThaispecies Angustopilaelevata (ThompsonetUpatham, 1997).