Syzygium hancei Merr. & L.M.Perry

Soh, Wuu-Kuang & Parnell, John, 2015, A revision of Syzygium Gaertn. (Myrtaceae) in Indochina (Cambodia, Laos and Vietnam), Adansonia 37 (2), pp. 179-275 : 228-230

28. Syzygium hancei Merr. & L.M.Perry ( Figs 23 ; 24 )

Journal of the Arnold Arboretum 19: 242 (1938); P.H.Hô, An Illustrated Flora of Vietnam (Câyco Viêtnam) 2: 51, fig. 3750 (1992); K.D.Nguyen, ( Myrtaceae ) Checklist of Plant Species of Vietnam 2: 904 (2003); J. Chen & Craven, Flora of China 13: 356 (2007). — Eugenia minutiflora Hance , Journal of Botany, British and Foreign 9: 5 (1871); non Syzygium minutiflorum (Bedd.) Gamble (1919) . — Type: China, Kwangtung, Hui-tung, Sampton & Hance 13754 (holo-, BM!; iso-, A![frag., A00069409], K!). (see Note 1)

Eugenia bonii Gagnep., Notulae Systematicae 3: 318. 1917; Gagnep., in Lecomte, Flore générale de l’Indochine 2: 808, fig. 87 (1920). — Syzygium bonii (Gagnep.) Merr. & L.M.Perry , Journal of the Arnold Arboretum 19: 102 (1938); P.H.Hô, An Illustrated Flora of Vietnam (Câyco Viêtnam) 2: 48, fig. 3729 (1992); K.D.Nguyen, ( Myrtaceae ) Checklist of Plant Species of Vietnam 2: 901 (2003). — Type: Vietnam [ Annam], Than-hoa, Cua-bang, fl., 17.VIII.1812, Bon 5603 (lecto-, P![P00589297], here designated; isolecto-, P![2 sheets, one sheet with barcode P00589368 & the other sheet without barcode and with Gagnepain’s drawing attached]), syn. nov.

Eugenia campylocarpa Gagnep., Notulae Systematicae (Paris) 3: 320 (1917). — Syzygium campylocarpum (Gagnep.) Merr. & L.M.Perry , Journal of the Arnold Arboretum 19: 116 (1938). — Type: Laos, Attopeu, Bolaven plateau [Attopeu plateau], fl., III.1877 Harmand 1257 (lecto-, P![P00589291], here designated; isolecto-, P![4 sheets: 2 sheets with barcodesP0058992 &P00589293, both with tickets bearing Harmand’s signature, 2 other sheets without barcode, Pierre 3189 [Harmand 1257]), syn. nov. (see Note 2).

Eugenia mekongensis Gagnep., Notulae Systematicae 3: 328 (1918); Gagnep. in Lecomte, Flore générale de l’Indochine 2: 807 (1920). — Syzygium mekongense (Gagnep.) Merr. & L.M.Perry , Journal of the Arnold Arboretum 19: 102 (1938); P.H.Hô, An Illustrated Flora of Vietnam (Câyco Viêtnam) 2: 54, fig. 3747 (1992); K.D.Nguyen, ( Myrtaceae ) Checklist of Plant Species of Vietnam 2: 906 (2003); D.D.Soejarto et al. Seed Plants of Cuc Phuong National Park: 432 (2004), [excl. S. szemoaense = Memecylon sp. ]; M.F.Newman et al., A Checklist of the Vascular Plants of Lao PDR: 245 (2007). — Type: Thailand, between Ubon and Kemmarat, Ubon river, fl., Thorel s.n. (lecto-, P![P00589243], here designated). — Remaining former syntypes: Cambodia, Kandal, around Phnom Penh, Gourgand (A n.v., P!); Vietnam [ Annam], Than-hoa, fr., Bon 5808 (P![2 sheets, P0089195 & P00589196]); Vietnam [ Cochinchina], Dong Nai, Baria, Mt. Dinh, Pierre s.n. (P!), syn. nov.

Syzygium petelotii Merr. & L.M.Perry , Journal of the Arnold Arboretum 19: 103 (1938); P.H.Hô, An Illustrated Flora of Vietnam (Câyco Viêtnam) 2: 55, fig. 3752 (1992); K.D.Nguyen, ( Myrtaceae ) Checklist of Plant Species of Vietnam 2: 907 (2003). — Type: Vietnam [ Tonkin], Phu Tho, Phu Ho, Pételot 1043 (holo-, A!; US!), syn. nov.

Syzygium szemaoense Merr. & L.M.Perry , Journal of the Arnold Arboretum 19: 105 (1938); P.H.Hô, An Illustrated Flora of Vietnam (CâycoViêtnam) 2: 61, fig. 3770 (1992); K.D.Nguyen, ( Myrtaceae ) Checklist of Plant Species of Vietnam 2: 908 (2003); J. Chen & Craven, Flora of China 13: 352 (2007). — Type: China, Yunnan, Szemao, Henry 12138 (holo-, NY!; iso-, A!, K!), syn. nov.

ADDITIONAL MATERIAL EXAMINED. — Cambodia. Kampong Spoe: Phnum Aural, 8.III.2001, K. Eanghourt, M.Sophal & S.Kosal MoE Herb. Cat. 670 ( K). — Kampot: Mt. Bokor , Popok Vil waterfall, 8.III.2001, Middleton & M.Monyrak 623 (P); idem, 11.XII.2000, S.Hul et al. 954 (P). — Kandal: Phnom Penh, Gourgand s.n. (P). — sin. loc. Gourgand s.n. (P) .

Vietnam. Cochinchina: X.1866, Pierre s.n. (P). — Da Nang: Son Tra , 9.VII.1986, Anon. LX-VN 2768 ( HN) . — Dac Lak: Duc Minh , 9.XII.1979, Anon. 574 ( HN) ; Dak Nang, Dao Nghia , 17.V.1979, Phuong 925 ( HN) . — Dong Nai: Mt. Dinh, Pierre s.n. ( BM, E). — Gia Lai: Ayun Pa , 21.IV.1988, Bien 764 ( HN) ; K’bang, 20.XI.2005, N.Q. Binh & D.D.Cuong VN 1620 ( HN) . — Ha Giang: Bac Quong , 14.VII.2001, N.Q.Binh, D.D.Cuong & Schnitzler VN 848 ( HN) . — Ha Tay: Mt. Ba Vi, Pételot 2596 (A). — Khanh Hoa: Nha Trang , VI.1909, d’Alleizette s.n. (L). — Kon Tum: 30.XI.1978, H.H.Dung 383 ( HN) . — Lam Dong: Bao Loc, Lien 252 ( HN) ; Da Chay, 16.IV.1997, Averyanov & N.Q.Binh VH 3988 ( HN) ; Daa Lao, 16.V.1953, Schmid s.n. (P). — Lao Cai: Chapa, VII.1931, Pételot 4678 (A). — Ninh Binh: Nho Quan, Cuc Phuong National Park , 8.VIII.1999, N.M.Cuong, H.L.Quyen & M.V.Xinh NMC 402 ( CPNP) ; Nho Quan, Cuc Phuong National Park , 17.IX.1999, N.M.Cuong, H.L.Quyen & M.V. Xinh NMC 523 ( CPNP) ; Nho Quan, Cuc Phuong National Park , 24.VII.2000, N.M.Cuong, H.L.Quyen & M.V. Xinh NMC 877 ( CPNP) . — Quang Ninh: Ha Coi, Tsai Wong Mo Shan and vicinity, XI.1936, W.T.Tsang 27247 ( K, P) ; Tien Yen, Kau Nga Shan and vicinity, I.1937, W.T.Tsang 27485 ( K, P) ; idem, XII.1936, W.T. Tsang 27389 ( K, P) ; Dam Ha , Sai Vong Mo Leng, W.T. Tsang 30068 (E, P) ; idem, VII.1940, W.T. Tsang 30383 ( K, P) ; idem, VII.1940, W.T. Tsang 30417 (A, K, P) ;. — Than-hoa: Bon 5808 (P). — Thua Thien-Hue: Bach Ma, Nham 559 ( HN) . — Vinh Phuc: Me Linh, Ngoc Thanh , 27.VI.2000, Phuong 3049 ( HN) .

DISTRIBUTION. — China (Fujian, Guangdong, Guangxi and Hainan), Cambodia, Laos, Thailand and Vietnam ( Fig. 27 ).

ECOLOGY OR HABITAT. — Primary and secondary forest, and along rivers.

CONSERVATION STATUS. — IUCN Global Status: Least Concern ( LC); IUCN Regional (Indochina) Status: Least Concern ( LC).

PHENOLOGY. — Flowering: March to October. Fruiting: November to April.

VERNACULAR NAMES. — Cambodia: Hong tuk, Pring teuk (Kampong Spoe). Vietnam: Phen den, Tram, Tram cuulong.

USES. — The fruit is edible.

DESCRIPTION

Shrub or tree to 20 m tall, to 25 cm dbh. Bark brownish or greyish, slash bark dark purplish. Glabrous.

Figures

FIG. 23. — Holotype of Syzygium hancei Merr. & L.M.Perry, Sampton & Hance 13754 (BM).

FIG. 24. — Lectotype of Syzygium mekongense (Gagnep.) Merr. & L.M.Perry, Thorel s.n. (P00589243).

FIG. 27. — Distribution of Syzygium hancei Merr. & L.M.Perry (), Syzygium harmandii (Gagnep.) Merr.& L.M.Perry () and Syzygium hemisphericum (Wight) Alston () in Indochina.