Paramaya De Haan, 1837

Ng, Peter K. L. & Forges, Bertrand Richer De, 2015, Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific, Raffles Bulletin of Zoology 63, pp. 110-225 : 155

Key to species of Paramaya De Haan, 1837

1. Pseudorostral horns relatively short; ambulatory meri in adults relatively stout, short ( Fig. 21A–D ); G1 gently curved ( Fig. 23A–C ); Japan and Taiwan...................................................... ..................................... Paramaya spinigera (De Haan, 1837)

– Pseudorostral horns very long; ambulatory meri in adults relatively slender, long ( Figs. 21E, F , 22A ); G1 otherwise ..2

2. Surface of thoracic sternum relatively smooth, with very low rounded granules ( Fig. 50C ); G1 gently curved ( Fig. 23G–I ); Vanuatu........................................... Paramaya coccinea n. sp.

– Surface of thoracic sternum distinctly granulated ( Fig. 50B ); G1 more strongly curved especially distally ( Fig. 23D–F ); Philippines............................................. Paramaya ouch n. sp.

Figures

Fig. 21. General habitus, Paramaya species. A, P. spinigera (as Maja (Paramaya) spinigera, after De Haan, 1837: pl. 24, fig. 4); B, P. spinigera, dried female (70.1 × 58.8 mm) (KPM NH4195), Japan; C, P. spinigera, male (64.7 × 51.2 mm) (NHM 1961.6.5.117), Japan; D, P. spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; E, P. ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; F, P. ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines.

Fig. 23. Left G1s, Paramaya species. A–C, P. spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; D–F, P. ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; G–I, P. coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu. Scale bars = 5.0 mm [A, D, G]; 1.0 mm [B, C, E, F, H, I].

Fig. 22. General habitus. A, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; B, “Maja spinigera” (ca. 70 mm carapace length), Sri Lanka, not preserved (photograph: R. Pethiyagoda); C, D, “Maja spinigera” (after Alcock & Anderson, 1898: pl. 34, fig. 3).

Fig. 50. Male anterior thoracic sternums and abdomens. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea.