Tomosvaryella docta De Meyer, 1995

Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott, 2021, Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species, Zootaxa 5002 (1), pp. 1-103 : 25

Tomosvaryella docta De Meyer, 1995

Figs 11A–F , 52

Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, both broadened in basal half; right surstylus narrowed moderately in in apical half, left surstylus narrowed abruptly in apical half ( Fig. 11A ); epandrium longer than wide ( Fig. 11A ); subepandrial sclerite wide at the base in ventral view ( Fig. 11B ); gonopods with an unequal projection, right is farther than left one ( Fig. 11B ); phallus trifid, one with some small teeth-like projections in lateral view ( Fig. 11F ).

Specimens examined: IRAN: Khuzestan: Shush, 32°06’N, 48°26’E, 55 m a.s.l., 29–31.viii.2013, M. Parchami-Araghi, Malaise trap, JSS52189 (1♀, CNC); Sistan & Baluchestan: Daman, 27°24’N, 60°49’E, 2–12.vii.2016, H. Davari, Malaise trap, JSS51822 (1♂, CNC); Saravan, 27°25’N, 62°17’E, 17.x.2016, F. Hamzavi, sweep net, JSS51827 (1♀, CNC); Zabol, Dostmohammad, 31°09’N, 61°43’E, 470 m a.s.l., 29.vi.2016, M. Enaiatnia, Mal- aise trap, JSS51933 (1♀, CNC); Zabol, 31° 0 3’N, 61°22’E, 480 m a.s.l., 10.vii.2016, M. Enaiatnia, Malaise trap, JSS51860 (1♂, CNC); 31°02’N, 61°32’E, 470 m a.s.l., 2.ix.2016, M. Enaiatnia, Malaise trap, JSS51907 (1♀, CNC); Kerman: Fariab, Horesfandi, 28°08’N, 57°16’E, 5.viii.2016, M. Arabzadeh, Malaise trap, JSS51922 (1♂, CNC); ISRAEL: Hazeva Field School, 30°43’N, 35°15’E, 2.x.1993, S. Plotkin, Malaise trap, JSS51704 (1♂, TAU); 5.x.1997, S. Plotkin, Malaise trap, JSS52080 (1♂, TAU); Hazeva, Field School, 30°43’N, 35°15’E, 11.x.1997, A. Maklakov, Malaise trap, JSS51769–70 (2♂, TAU); CYPRUS: Kyrenia, 35°21’N, 33°09’E, 10–17.ix.2017, O. Ozden, Malaise trap, JSS52344 (1♂, CNC); 16–30.vii.2017, O. Ozden, Malaise trap, JSS52325, JSS52355 (2♂, CNC); 24.ix–1.x.2017, O. Ozden, Malaise trap, JSS52354 (1♂, CNC); 30.vii–13.viii.2017, O. Ozden, Malaise trap, JSS52366 (1♂, CNC); 9–16.vii.2017, O. Ozden, Malaise trap, JSS52330 (1♂, CNC); UNITED ARAB EMIR- ATES: Abu Dhabi: Al Wathba Wetland Reserve, 24°15’N, 54°37’E, i.2015, A. Saji & A. van Harten, Malaise trap, CNC470746 (1♂, CNC); ii.2015, A. Saji & A. van Harten, Malaise trap, CNC470512, CNC470572 (2♂, USNM); iii.2015, A. Saji & A. van Harten, Malaise trap, CNC470645, CNC470654, CNC470679–80 (4♂, CNC); iv.2013, A. Saji & A. van Harten, Malaise trap, CNC470729 (1♂, CNC); iv.2015, A. Saji & A. van Harten, Mal- aise trap, CNC470808 (1♂, CNC); ix.2013, A. Saji & A. van Harten, Malaise trap, CNC465503, CNC470708–10, CNC470713, CNC470716 (6♂, CNC); xi.2013, A. Saji & A. van Harten, Malaise trap, CNC470721, CNC470725, CNC470728 (3♂, CNC); Al Bida’a Protected Area, 23°36’N, 53°58’E, 29.x.2018, A. Saji & A. van Harten, Malaise trap, JSS52267–70 (4♂, CNC); Al Tawi Protected Area, 23°50’N, 53°10’E, 1.xi.2018, A. Saji & A. van Harten, Malaise trap, JSS52295–98 (4♂, CNC); Barqa Al Suqoor Protected Area, 23°60’N, 52°41’E, 20.iii.2018, A. Saji & A. van Harten, Malaise trap, JSS52260 (1♂, CNC); 29.x.2018, A. Saji & A. van Harten, Malaise trap, JSS52282 (1♂, CNC); Wadi Wurayah farm, 25°14’N, 56°11’E, 17–24.iii.2009, A. van Harten, Malaise trap, CNCD137123 (1♂, CNC); 19.iv–19.v.2009, A. van Harten, Malaise trap, CNCD137544 (1♂, CNC); 31.v–14.vi.2009, A. van Harten, Malaise trap, CNCD137579, CNCD137582 (2♂, EAD); Wadi Wurayah, 25°14’N, 56°11’E, 19–26.iv.2009, A. van Harten, Malaise trap, CNCD160581 (1♂, EAD).

Distribution: Cyprus, Egypt, Iran, Israel, Spain, United Arab Emirates ( Skevington 2020) ( Fig. 52 ).

Note: DNA barcodes of this species are genetically most similar to T. gazliensis , differing by 7.4% (pairwise divergence) (Supplementary Table 1).

Figures

FIGURE 11. Male genitalia of Tomosvaryella docta (CNCD137579, EAD) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view, F) phallus and phallic guide in lateral view.

FIGURE 52. Distribution of Tomosvaryella coquilletti, T. debruyni, T. docta in the Middle East.