Odochilus (Parodochilus) shavrini Rakovič & Anichtchenko, 2021

Rakovič, Miloslav & Anichtchenko, Alexander, 2021, Two new species of the genus Odochilus Harold, 1877 (Coleoptera: Scarabaeidae Aphodiinae: Odochilini), Zootaxa 4942 (3), pp. 447-450 : 447-449

Odochilus (Parodochilus) shavrini Rakovič & Anichtchenko , new species

( Figs. 1–3 )

Type material. Holotype, male: “ Philippines, Mindanao , Maragusan , 700m, Marangig Falls, Candalaga Mts., 7˚20’32.2”N 126˚10’24.0”E, 3- 4.05.2019, Shavrin A. V. leg.” ( DUBC) . Paratype, female, with the same label data ( DUBC) .

Description of the holotype. Small (length of 2.8 mm), strongly convex ( Figs. 1, 2 ), remarkably broader behind, length-to-width ratio of 1.98, dark brown, tibiae and tarsi light brown, antennae and mouth parts yellowish.

Head glabrous from above, except for minute, dust-like setae along ridges above eyes. Genae moderately protruding. Clypeus margin with upper and lower edges: upper edge convex, sharp and distinct, lower edge obsolete and not observable from above. Clypeal disc convex, matte, with rather uneven, corrugately punctate surface; longitudinal ridges on clypeus barely visible. Vertex with two pairs of distinct oblique ridges. Median pair of ridges replaced by a continuous semicircular ridge. Exterior pair forming a characteristic T-shaped structure with ridge extending forward from eyes. Eyes visible from above.

Pronotum transverse, 1.5 times as wide as long; opaque, convex, with four transverse ridges. Anterior and lateral margins, and all ridges with very small, dust-like setae. Lateral margins with two sub-angular emarginations and thus, with three moderately protruding, widely angular lobes (ponotum narrowed behind posterior lobe). Basal margin with apically slightly dilated setae. First and second pronotal ridges continuous, sharp, third ridge weakly and narrowly interrupted in the middle; fourth ridge broken into six discrete teeth; lateral longitudinal ridges small, connected with ends of third ridge. Intervals between ridges matte.

Elytra widely suboval, 1.1 times as long as wide; with small humeral tooth at base of costa 8. Odd costae twice as high as even ones; both odd and even costae covered by small, dust-like setae. Intercostae with large punctures; the punctures about as long as wide, distance between punctures slightly longer than their diameter. Intervals between punctures shiny. Scutellum small, elongate, with rounded apex.

Profemur wide, with large punctures throughout. Protibia slim at base; slightly widened toward the apex, with three equidistant teeth in apical half; apical tooth long, medial slightly shorter, basal tooth twice shorter than medial. Apical spur as long as two basal tarsomeres.

Ventral side ( Fig. 3 ) dark brown, matte. Abdomen with ventrites 1–5 minutely punctate, ventrite 5 with four short setae; ventrite 6 densely punctate, matte, with six setae. Pygidium with six erect setae.

Female: there are no considerable differences between external characters of the male holotype and those of the female paratype. The body length of the female paratype (3.0 mm) is larger compared to the male holotype (2.8 mm).

Differential diagnosis. The new species can be compared only with Odochilus (Parodochilus) buruensis Boucomont, 1926 , known from the Moluccas, because of an unusually small elytra length-to-width ratio (the elytra are relatively very broad). The two species can be most easily differentiated from each other based on the pronotal sculpture. In O. (P.) buruensis ridges 1–3 are in the form of continuous, high, narrow and sharp lamellae ( Fig. 4 ); in the new species, ridges 1–3 are convex but not in the form of lamellae and ridge 3 is rather interrupted medially ( Fig. 5 ). Longitudinal V-shaped formations, present in the subgenus Parodochilus instead of ridge 4, are very distinct in the former species ( Fig. 4 ), but rather indistinct in the new species ( Fig. 5 ).

Distribution. Species known only from the type locality in Mindanao Island, The Philippines.

Name derivation. New species named after our friend and colleague, specialist in Staphylinidae (Coleoptera) , Dr. Alexey Shavrin, who collected the new species.

Figures

FIGURES 1–6. Habitus of Odochilus (Parodochilus) species. 1, O. (P.) shavrini Rakovič & Anichtchenko, new species, male (holotype), habitus in dorsal view; 2, O. (P.) shavrini new species, male (holotype), habitus in dorsolateral view; 3, O. (P.) shavrini new species, male (holotype), habitus in ventral view; 4, O. (P.) buruensis Boucomont, 1926, drawing of pronotum overprinted from Rakovič, 1997 (permitted by Acta Societatis Zoologicae Bohemicae); 5, O. (P.) borneensis Rakovič & Anich- tchenko, new species, female (holotype), habitus in dorsal view; 6, O. (P.) jani Bezděk & Křivan, 2001, paratype no. 3, habitus in dorsal view.