Zyras (Zyras) hastatus FAUVEL, 1904

Assing, Volker, 2017, On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini), Beiträge Zur Entomologie = Contributions to Entomology 67 (1), pp. 117-192 : 139

Zyras (Zyras) hastatus FAUVEL, 1904

( Figs 16 , 54 , 90 , Map 7 )

Zyras hastatus FAUVEL, 1904: 64 .

Type material examined: Syntypes: 1 ♀: “ Belgaum / hastata Fvl. / Coll. I.R.Sc.N.B. / Syntypus Zyras hastatus Fauvel , rev. V. Assing 2016” ( IRSNB); 1 ♀ [damaged; both antennae missing]: “Cotype / Belgaum / 1266 / hastata Fvl. / Syntypus ♀ Zyras hastatus Fauvel , rev. V. Assing 2016” ( BMNH) .

Comment: The original description is based on at least two syntypes (“Sexus differentia latet”) from “Belgaum” ( FAUVEL 1904). One syntype was located in the collections of the IRSNB and another in the collections of the BMNH. Both syntypes are females.

Redescription: Body length 4.1–6.0 mm; length of forebody 2.1–2.8 mm. Coloration ( Figs 16 , 54 , 90 ): head black; pronotum pale-reddish; elytra reddish-yellow with the postero-lateral portion extensively blackish (leaving only the scutellum and its vicinity, the anterior margin, and the suture reddish-yellow); abdomen bicoloured with tergites II–V pale-reddish, tergite VI black with the anterior margin and the antero-lateral portions reddish, and tergites VII–VIII black; legs pale-yellowish; antennae dark-brown with antennomeres I–III dark-reddish and the apical half of antennomere XI dark-reddish; maxillary palpi reddish with the terminal palpomere yellowish.

Head ( Fig. 54 ) moderately transverse, median portion extensively impunctate; punctures in lateral portions moderately dense and large, but shallow. Eyes moderately large, but weakly convex, much longer than postocular region in dorsal view. Antenna ( Fig. 16 ) 1.75 mm long; antennomeres IV approximately as long as broad, V weakly transverse, VI–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI conspicuously elongate, nearly as long as the combined length of VII–X.

Pronotum ( Fig. 54 ) distinctly transverse, approximately 1.2 times as broad as long and 1.3 times as broad as head, broadest anterior to middle; lateral margins smoothly convex (dorsal view); punctation not very coarse, rather sparse, and somewhat irregularly distributed; midline narrowly impunctate; lateral margins each with four long, black setae, anterior margin with an additional long black seta on either side.

Elytra ( Fig. 54 ) 0.85 times as long as pronotum; punctation moderately coarse, defined, rather dense, and regularly distributed; interstices on average distinctly broader than diameter of punctures. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of II–IV.

Abdomen ( Fig. 90 ) narrower than elytra, with deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of non-setiferous punctures; tergite III with a lateral long brown seta on either side, posterior margin with three long brown setae on either side and with a median pair of fine short yellowish setae; tergites IV–V with a median pair of punctures, with a lateral seta on either side, posterior margins with four long brown setae on either side, and with a median pair of minute pale setae; tergite VI anteriorly only with few shallow non-setiferous punctures laterally (without such punctures in the middle), with a median pair of punctures, and with approximately 12 setiferous punctures at posterior margin, otherwise impunctate (except for scattered micropunctures); tergite VII anteriorly with a transverse row (not band!) of sparse non-setiferous punctures, with a transverse row of six setiferous punctures at posterior third, and with additional setiferous punctures at posterior margin; tergite VIII with rather sparse long setae only in posterior portion, posterior margin convex, in the middle with indistinct and small concavity.

♂: unknown.

Comparative notes: Among the species of similar coloration (e.g., Z. beijingensis PACE, 1993 ), Z. hastatus is characterized by slender antennae with a conspicuously elongate antennomere XI, a relatively strongly transverse pronotum with smoothly convex lateral margins, and the punctation pattern of the abdomen (particularly of tergites VI and VII).

Distribution: This species is currently known only from the type locality in South India ( Map 7 ).

Figures

Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions.

Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm.

Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm.

Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm.