Pachycephala tenebrosa Rothschild, 1911

Pachycephalidae, Aegithalidae, Remizidae, Paridae, Sittidae, Neosittidae, Certhiidae, Rhabdornithidae, Climacteridae, Dicaeidae, Pardalotidae, Nectariniidae, And & Lecroy, Mary, 2010, Type Specimens Of Birds In The American Museum Of Natural History Part 8. Passeriformes:, Bulletin of the American Museum of Natural History 2010 (333), pp. 1-178 : 11-12

Pachycephala tenebrosa Rothschild

Pachycephala tenebrosa Rothschild, 1911: 20 (Mount Goliath) .

Now Colluricincla tenebrosa tenebrosa (Rothschild, 1911) . See Coates, 1990: 220, Dickinson, 2003: 489, Boles, 2007: 432, and Dumbacher et al., 2008.

HOLOTYPE: AMNH 659480 , adult male, collected on Mt. Goliath , 04.40S, 139.52E ( USBGN, 1982), Oranje Mountains, Papua Province, Indonesia, on 15 February 1911, by Albert S. Meek (no. 5394). From the Rothschild Collection.

COMMENTS: Rothschild cited Meek’s unique field number of the holotype in the original description and noted that he had nine specimens. Paratypes are: AMNH 659477–659479 , 659481 , and 659482, males ; and AMNH 659483– 659485 , females, all collected on Mt. Goliath

in January and February 1911 by Meek, whose collection was reported on by Rothschild and Hartert (1913).

Sibley and Monroe (1990: 461) and Dickinson (2003: 489, citing Diamond, 1972), have put both Pachycephala tenebrosa Rothschild, 1911 , and Rhectes tenebrosus Hartlaub and Finsch, 1868 (listed as Pitohui tenebrosus by Mayr, 1967: 51), from Palau in the genus Colluricincla . When this is done, Rothschild’s name being the junior name, the next available name becomes Melanorhectes umbrinus Reichenow, 1915; and Pachycephala tenebrosa Rothschild is listed by Dickinson as Colluricincla umbrina (Reichenow, 1915) . This is followed by Boles (2007: 432). However, Diamond (1972: 274–275, 284) did not suggest putting Pachycephala tenebrosa in Colluricincla , only that Pitohui tenebrosus be shifted to Colluricincla .

Recent DNA analysis by Dumbacher et al. (2008) has shown quite clearly that Pitohui tenebrosus is closely related to Pachycephala pectoralis and should be included in Pachycephala , the name, confusingly, becoming Pachycephala tenebrosa (Hartlaub and Finsch) .

The suggestion that Pachycephala tenebrosa Rothschild, 1911 , be put in Colluricincla apparently dates from Sibley and Ahlquist (1982), where such a suggestion was made by Richard Schodde. Dumbacher et al. (2008) did not include Pachycephala tenebrosa Rothschild in their DNA analysis, but ongoing molecular phylogenetic investigation within Colluricincla confirms the placement in that genus of Pachycephala tenebrosa Rothschild through the inclusion of DNA samples from the Schrader Range and from Mt. Stolle (B. Benz, personal commun., 2010).