Gyrophaena (Gyrophaena) joyi Wendeler, 1924

Enushchenko, Ilya V. & Semenov, Viktor B., 2016, A review of the genus Gyrophaena Mannerheim 1830 (Coleoptera: Staphylinidae: Aleocharinae: Gyrophaenina) of the Caucasus and adjacent territories, Zootaxa 4126 (3), pp. 301-337 : 314-316

Gyrophaena (Gyrophaena) joyi Wendeler, 1924

(Figs. 25–27)

Gyrophaena joyi Wendeler, 1924: 344 ; Enushchenko & Shavrin, 2011: 1204 Gyrophaena convexicollis Joy, 1912: 148

Gyrophaena joyi asiatica Wüsthoff, 1937: 143

Gyrophaena joyi ; Nikitskiy & al., 2008: 112

Material examined. RUSSIA: STAVROPOL TERRITORY: Predgorniy District: 3 Ƌ, 4 ♀: Mineral'niye Vody, Beshtau Mountain, 4–17.VIII.1993, E.A. Khachikov leg. ( ZMM); KRASNODAR TERRITORY: Krymskiy District: 8 Ƌ, 7 ♀: Krymsk, right side of Adagum River, 5.X.2011, I.V. Enushchenko leg. (cIE, ZMM); 3 Ƌ, 6 ♀: ibid., on Fomes fomentarius , 5.X.2011, I.V. Enushchenko leg. (cIE, ZMM); Severskiy District: 1 Ƌ, 1 ♀: Ubinskaya, 21.VI.1975, V.V. Belov leg. ( ZMM); 7 spec.: ibid., VI.1986, I.A. Ushakov leg. (cVS); 7 spec.: environs of Ubinskaya, Papai Mountain, 2.VI.1986, I.A. Ushakov leg. (cVS); 9 spec.: ibid., Derbiy Mountain, 3.VI.1986, I.A. Ushakov leg. (cVS); Apsheronskiy District: 1 Ƌ: N slopes of Skalisty Mts. rng., near Peredovaya River 1.st, Sukhaya balka, sift, pitfall traps, 901 m, N 44°02ʹ46,1ʺ E 41°26ʹ04,2ʺ, 27.05.2014, A. Pütz leg. (cAP); 1 Ƌ, 2 ♀: valley of Kurdzhips River, Mezmai, 4–17.VII.1993, E.A. Khachikov leg. ( ZMM); Anapskiy District: 4 Ƌ, 6 ♀: 3 km E Bol'shoi Utrish, 24–30.VIII.2003, D. Popov, A. Rudaikov, E. Khachikov leg. ( ZMM); Khostinskiy District: 1 Ƌ: Maryino, 25.IX.1988, I.A. Ushakov leg. (cVS); 48 spec.: Khosta, 1–19.VI.1988, V.B. Semenov leg. (cVS); Adlerskiy District: 1 Ƌ, 2 ♀: 23 km NE Adler, lower course of Chvizhepse River, 43° 38' 25" N; 40° 04' 33" E, window trap, 28.IV–4.V.2013, K. Makarov leg. (cVS); KARACHAY- CHERKESSIA: Karachay District: 10 Ƌ, 14 ♀:

FIGURES 24–27.

Teberdinskiy Biosphere Reserve, Teberda, VII.2011, E.A. Khachikov leg. (cEKh, cIE, ZMM); 1 Ƌ, 1 ♀: ibid., Khatipara Mountain, 17.VII.2012, E.A. Khachikov leg. ( ZMM); NORTH OSSETIA: Alagirskiy District: 1 Ƌ: North- Ossetian Biosphere Reserve, Tseiskoye Gorge, Alnus -brushwoods, 20.VII.1982, S.K. Alexeev leg. (cVS); ABKHAZIA: Gagra District: 1 Ƌ: Avadkhara, 24.V.2004, E.A. Khachikov leg. ( ZMM); 2 Ƌ, 1 ♀: Mjusserskiy Biosphere Reserve, 18.IX.1988, I.A. Ushakov leg. (cVS); Gudautskiy District: 16 spec.: Duripsh, 10–16.IX.1985, I.A.Ushakov leg. (cVS); Sukhum District: 1 ♀: Gumistinskiy Biosphere Reserve, Tsumuri, 400–500 m, 4.VI.1982, Drabkin leg. ( ZIN); 1 Ƌ, 1 ♀: Kelasuri Canyon, 18.IX.1988, I.A. Ushakov leg. ( ZMM); Ochamchirskiy District: 7 Ƌ, 3 ♀: Dzhgerda, foothills of Kodorsky Ridge, 21.IX.1985, I.A. Ushakov leg (cVS); GEORGIA: SAMTSKHE- JAVAKHETI: 1 ♀: ‘Borshom, Kaukasus’ [‘ Gyrophaena fasciata ’] ( MAKB); KAKHETI: Sagaredzhskiy District: 1 ♀: Mariamdjvari Biosphere Reserve, Carpinus forest, 13–14.V.1987, S.I. Golovatch & K.Yu. Eskov leg. (cVS); ARMENIA: TAVUSH AREA: Shamshadyn District: 1 spec.: Dilizhan Biosphere Reserve, valley Akhnabat near Salakh, 1400–1500 m, Carpinus etc. forest, 17.IV.1989, S.I. Golovatch leg. (cVS); AZERBAIJAN: Astarinskiy District: 1 Ƌ, 1 ♀: Astara Istisu, W Astara, 100 m, 2–6.VI.1996, S.I. Golovatch leg. ( ZMM); 4 Ƌ, 9 ♀: Bilyasyar, 800ʹ on Vasharu-Chay River, 16.VII.1932, Znoiko leg. ( ZIN).

Additional material examined: RUSSIA: REPUBLIC OF CRIMEA: Bakhchisaraiskiy District: 1 Ƌ: Kokoz [Sokolinoye], 9.V. [19]11, V. Pliginskiy leg. ( ZIN); Yalta: 1 spec.: Angarskiy Pass, 28.V.1911, V. Pliginskiy leg. ( ZIN); 1 Ƌ: Taushan-Bazar, 26.VI. [19]07, V. Pliginski leg. [‘ Gyrophaena bihamata’] ( ZIN); 1 Ƌ: ibid., 27.VI. [19]07, V. Pliginskiy leg. ( ZIN); 1 Ƌ: ibid. 14.VII. [19]07, V. Pliginskiy ( ZIN); Alushta: 2 ♀: Kosʹmo- Damianovskiy Mon[astery], 23.VI. [19]08, V. Pliginski leg. ( ZIN); 7 Ƌ, 4 ♀: ibid., 18.VI. [19]11, V. Pliginski leg. ( ZIN); Kirovskiy District: 2 Ƌ, 1 ♀: Stariy Crym, mountain forest, 15–16.VIII.2005, R. Cibuļskis leg. ( DUBC); VOLGOGRAD AREA: Kalachevskiy District: 1 Ƌ, 3 ♀: Kalach-na-Donu, 17–18. IX.1997, K.A. Grebennikov leg. (cIE); ROSTOV AREA: Sholokhovskiy District: 1 Ƌ, 3 ♀: Veshenskaya, 17.VI.2002, E.A. Khachikov leg. ( ZMM); Ust'-Donetskiy District: 1 Ƌ: Krymskiy, 85 km E Rostov-na-Donu, Savateyevskaya balka, 17.V.2008, V.N. Prasolov leg. ( ZIN); Azovskiy District: 52 spec.: Rostov-na-Donu, 21. VI.2009, E.A. Khachikov leg. (cEKh, ZMM).

Distribution. Europe, Caucasus, Turkey ( Korge 1971), Iran ( Assing 2011a), Siberia ( Enushchenko & Shavrin 2011, 2012b).

Remarks. G. joyi has been recently recorded from the Caucasus (Khosta) by Nikitskiy & al. (2008).