Achmonia congoensis ( Bernhauer, 1932 ) Bernhauer, 1932

Janák, Jiří & Bordoni, Arnaldo, 2014, Revision of the genus Achmonia of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae), Zootaxa 3872 (3), pp. 257-274 : 271-272

Achmonia congoensis ( Bernhauer, 1932) , comb. nov.

( Figs. 6, 11 , 68 , 71–74 )

Eulissus congoensis Bernhauer, 1932: 144 ; Scheerpeltz, 1933: 1317; Herman, 2001: 3627.

Type locality. D. R. Congo, Haut-Uele, Yebo Moto.

Type material examined. Lectotype (by present designation) ♀: DEMOCRATIC REPUBLIC OF CONGO: “Musée du Congo / Haut-Uele: Yebo Moto/ VII.1926 / L. Burgeon”, “R. DET./ 1664”, “ Eulissus / congoensis / Bernh.” (hand of Bernhauer), ( MRAC). This specimen is chosen as lectotype and the labels: “ Lectotypus Eulissus congoensis Bernh., Janák & Bordoni des. 2014” and “ Achmonia congoensis (Bernh.) , Janák & Bordoni det. 2014” were attached to this specimen.

Additional material examined. BURKINA FASO: 1 ♂: Haute Volta, Bobo-Dloulasso, vii–viii.1964, R. Siffointe ( MRAC). CÔTE D’IVOIRE: 1 ♂, Bingerville, vii.1962, J. Decelle ( MRAC). DEMOCRATIC REPUBLIC OF CONGO: 1 ♂: Lac Albert, Kasenyi, H. J. Brédo 15.v.1935 ( MRAC); 1 ♀: Lac Albert: Kasenyi, 1.v.1935, H. J. Brédo ( MRAC); 1 ♀: Kasenyi, i.1939, P. Lefèvre ( MRAC); 1 ♂, 1 ♀: P. N. G., Miss. H. De Saeger, II/ec/4, 5.iii.1951, Réc. H. De Saeger ( JJRC, MRAC); 1 ♂, P. N. G., Miss. H. De Saeger, II/me/4, 9.ii.1951, Réc. J. Verschuren ( MRAC). ETHIOPIA: 1 ♂: N. O. Africa, Süd. Aethiop., Malo Fl., O. Neumann S. V. ( JJRC). GHANA: 1 ♂: Northern Prov., Mole N. P., 18.vi.2006, S. Pokorný lgt. ( JJRC). GUINEA: 1 ♂: Guinée, coll. Fauvel ( IRSNB). TOGO: 1 ♀, Bismarkburg, L. Conradt S. ( MFNB).

Description. Body length ( Fig. 6 ) 13–19 mm; length from anterior margin of head to posterior margin of elytra: 8–10 mm. Head and pronotum black, elytra yellowish, scutellum and abdomen brown with slight bronze reflection, paratergites, sternites 3–5 and apical half of genital segment yellowish; antennae and anterior tarsi and tibiae reddish brown, middle and posterior tarsi and tibiae yellowish. Labrum quadrilobate, with only slightly prominent inner lobes ( Fig. 68 ). Head and pronotum with very fine micro-punctation. Head without postocular punctures ( Fig. 68 ), punctures near posterior margin with light long setae. Protrusion near posterior angles of head distinct. Temples with variable number of fine punctures, from almost impunctate to about 30 punctures. Anterior margin of pronotum with several fine, long setae. Elytra as long as pronotum, posteriad, slightly dilated and slightly wider than pronotum, with rounded humeral angles. Surface with fine and sparse punctation, arranged in three series; with additional punctures near posterior margin and some of these with very long, fine setae. Abdomen with very fine and dense, transverse micro-waves and fine and not sparse punctation.

Male. Temples sparsely to densely punctured (from about 5 to 30 fine punctures). Tergite and sternite of the male genital segment as in Figs. 73, 74 . Aedeagus length ( Fig. 71 ) 1.60–1.85 mm; median lobe very long, with enlarged apex; parameres very large ( Fig. 72 ); internal sac very narrow, covered by very fine and sparse scales.

Female. Temples smooth or at most sparsely punctured.

Ratios (n = 6): HL/HW: 0.82–0.87 (M 0.84); TeL/EyL: 1.71–1.92 (M 1.84); PL/PW: 0.98–1.06 (1.01); EL/EW: 0.86–0.96 (M 0.93); PW/HW: 0.88–0.98 (M 0.92); EW/PW: 1.02–1.09 (M 1.06); EL/PL: 0.95–0.99 (M 0.97); A5L/A5W: 0.50–0.57 (M 0.59); A10L/A10W: 0.50–0.62 (M 0.56).

Differential diagnosis. Achmonia congoensis differs from other Afrotropical species of the genus by the combination of brown, abdominal tergites with bronze reflections, yellowish pleurites and sternites 3–5, and by the shape of the aedeagus with a very long median lobe, even more enlarged apically then in A. amabilis .

Distribution. The species is distributed in Burkina Faso, Côte d’Ivoire, D. R. Congo, Ethiopia, Ghana, Guinea and Togo ( Fig. 2).

Remarks. As the labrum of Achmonia congoensis is dark, completely chitinised and quadrilobate ( Fig. 68 ), the species was transferred from Eulissus to Achmonia . In his description, Bernhauer (1932) mentioned the type localities Haut-Uele: Yebo Moto, and Tora. No specimens from Tora were available for study.

Figures

FIGURES 3 – 12. 3, 8, 12, Achmonia carinata sp. nov., HT male; 4, 7, 9, A. simulator sp. nov., PT male, 4, 9 from Mposhi, 7 from Joko; 5, 10, A. amabilis (Boheman), female from Ithala; 6, 11, A. congoensis (Bernh.), male from P. N. Garamba. 3 – 6, habitus; 7, lateral part of pronotum, ventral; 8 – 11, head; 12, pronotum and elytra. SL, superior line; IL, inferior line; EL, elytral length; p, postocular puncture. Scales 1 mm (3 – 6) and 0.5 mm (7 – 12).

FIGURES 54 – 68. 54 – 63, Achmonia simulator sp. nov., 54 – 57, 63, PT male from Joko, 58 – 61, HT male, 62, PT from Bolama; 64 – 67, A. amabilis (Boheman), 64, 66, 67, LT male, 65, female from Ithala; 68, A. congoensis (Bernh.), male from Malo. 54, aedeagus lateral; 55, 58, 61, aedeagus dorsal; 56, 59, paramere; 57, 60, median lobe of aedeagus; 62, sternite 9 and sclerites of tergite 10; 63, left metatibia and metatarsus; 64, 68, head; 66, sternite 9; 67, tergite 10; 65, lateral part of head. Scales 1 mm (64 = 65, 68), 0.5 mm (54 = 55, 58, 61, 62; 63; 66 = 67) and 0.1 mm (56 = 57, 59, 60).

FIGURES 69 – 74. 69 – 70, A. amabilis (Boheman), LT male; 71 – 74, A. congoensis (Bernh.), male from Malo. 69 – 70, aedeagus dorsal; 70, 72, paramere; 73, sternite 9; 74, tergite 10. Scale 0.5 mm (69 = 71, 73, 74; 70 = 72).