Eudorylas discretus ( Hardy, 1952 )

FÖLDVÁRI, MIHÁLY, 2013, Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae), Zootaxa 3656 (1), pp. 1-121 : 55

Eudorylas discretus ( Hardy, 1952)

( Figs 18A–G )

Dorilas (Eudorylas) discretus Hardy 1952: 9 .

Diagnosis: Third antennal segment long acuminate. Tips of surstyli turned towards ventral side (90°), IS broad, right side pale, probably thin, at the base finger-like protuberance on ventral side (OS as well); hypandrium peculiar shape, one lobe in the middle with striped membranous structures; phallic guide tube-like, with tooth-like structure facing SS in lateral view; phallus trifid, long and coiled.

Type material: Burundi: 1♂, HT, Urundi , Bururi, 13-iii-1949, alt. 1950 m, F. François, No 19.700 or 29.700. ( ISNB) .

Male

Head. Third antennal segment long acuminate (projection as long as 3rd segment); brown. Face silvery shining. Frons, upper part brown to black, lower part silvery shining; eyes touching for distance equal to 4 times ocellar triangle. Occiput, upper half brownish pollinose, lower half silver-brown pollinose.

Thorax. Humeri pale brown. Mesonotum (viewed obliquely from front) brownish pollinose; greyish from the side. Scutellum brownish pollinose, with weakly developed hairs. Dorsocentral hairs also weak. Halter yellowish brown.

Legs. Trochanters and base of femora yellow, femora brown, knees, tibiae yellow, hind tibia slightly curved. Tarsal segments brownish, last segment darker. Ventroapical row of 6–8 short spines on first four femora; no spines on 3rd femur. Subapical (distal) spines on first four tibiae distinct. No anteromedial hairs on 3rd tibia. Hind trochanter with smooth, whitish hairs on ventral side. Pulvilli shorter than last tarsal segment.

Wing. Fourth costal section 1.5 times as long as third costal section. Cross-vein R-M at 2/5 of discal cell. Pterostigma fully coloured. Hairs on tegula uncertain.

Abdomen. Viewed obliquely from front tergites brownish pollinose, hind margins and sides greyish. Hairs dispersed, short, whitish and weakly developed. 4 strong, black lateral bristles on first tergite. T5 slightly shorter than ST8. Genitalia without dissection: yellow, thickened.

Genitalia. Surstyli, tips turned towards ventral side (90°), IS broad, right side pale, probably thin, at the base fingerlike protuberance on ventral side (OS as well); epandrium distinctly smaller than ST8, broad; ST8 not extending to right side, ridge in the middle ( Fig. 18B ). SES very faint, in ventral view joins to GP; gonopods asymmetrical, right side longer, pointed in ventral view; hypandrium peculiar shape, one lobe in the middle with striped membranous structures; phallic guide broad, sides parallel, tube-like in ventral view ( Fig. 18F ); tooth-like structure facing SS in lateral view ( Fig. 18A ); phallus trifid, branches close to each other, long, coiled; ejaculatory apodeme 3-sided; sperm pump with two flattened projections, otherwise rounded, elliptic ( Fig. 18D ).

Female—Unknown.

Remarks— It is not clear how the projection attaches to IS at the basal part in lateral view.

Distribution— Burundi.

Figures

FIGURE 18. Male genitalia of Eudorylas discretus (A–G) and E. diversus (H–M).—A: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: sperm pump with ejaculatory apodeme, E: surstyli, dorsal view, F: ventral view with phallic guide and phallus (without ST8), G: sperm pump with ejaculatory apodeme, H: outer surstylus with phallic guide and phallus, lateral view, I: dorsal view, J: inner surstylus, lateral view, K: sperm pump with ejaculatory apodeme, L: surstyli, dorsal view, M: ventral view with phallic guide and phallus (without ST8). Scale bars: 0.1 mm