Helix (Pelasga) figulina ROSSMÄSSLER, 1839

Neubert, Eike, 2014, Revision of Helix LINNAEUS, 1758 in its eastern Mediterranean distribution area, and reassignment of Helix godetiana KOBELT, 1878 to Maltzanella HESSE, 1917 (Gastropoda, Pulmonata, Helicidae), Contributions to Natural History 26, pp. 1-200 : 144-153

Helix (Pelasga) figulina ROSSMÄSSLER, 1839 ( Figs 227–233 )

1839 Helix figulina ROSSMÄSSLER, Iconographie der Land- und Süsswassermollusken (1), 2 (3/4): 9, Taf. 44, Fig. 580 [ Griechenland].

1904 Helix (Helicogena) pomacella var. attalus KOBELT, Iconographie der Land- und Süsswassermollusken 2 (10): 70, Taf. 299, Fig. 1917 [Pergamon].

1904 Helix (Helicogena) pelasgica KOBELT, Systematisches Conchylien-Cabinet VI : 192, Taf. 345, Fig. 6–11 [Phthiotis, Hypati].

Type specimens:

figulina : holotype SMF 9845: H = 22.6, D = 22.1, PH = 17.2, PD = 11.6, PrD = 3.5, W = 4.75. pelasgica : lectotype SMF 9706 (SD Zilch 1952: 138): H = 32.6, D = 28.4, PH = 22.7, PD = 16, PrD = 3.4, W = 5.5; paralectotype pelasgica SMF 9708: H = 30.1, D = 31.2, PH = 23.5, PD = 16.8, PrD = 3.3, W = 5.25. attalus: lectotype SMF 9846: H = 21.2, D = 20.7, PH = 15.7, PD = 10.8, PrD = 3.8, W = 4.75.

Specimens examined:

Bulgaria: SMF 18529, Tripolitza (not identified, probably Tripolitza on the Peloponnese?).

FYROM: SMF 74203/3, between Skopje and Zelenikovo, 41.9022 21.5456, coll. K.L. Pfeiffer ex Knipper *; Skopje , 10 Km E Skopje, 41.9872 21. 6565, 293 m alt., 11.7.1972, NMBE 524571 /2; SMF 74201/2 Skopje 4 km N of the city, 42.0482 21.4206, coll. K.L. Pfeiffer ex Knipper*; SMF 311935/13 at the Vardar, ex RÖmheld 1916 *.

Greece: SMF 24475 unknown (probably Greece, probable typoid specimen of figulina ?), coll. Rossmässler; SMF 299394/7+2, Greece, coll. Krüper. Makedonia: Kozani, Kila close Kozani , 40.3306 21.7886, 21.06.1985, HAUS (1 alk.) ; Limni Pikrolimni, Mikrokambos , 40.85 22.82, 50 m alt., 10.5.2003, NMBE 19277 / 2; Kozani, Servia 6.5 km in dir. Kozani , 40.2398 21.9661, 21.06.1985, HAUS (1) ; Kozani , dry rocky slope 2 km S of Kozani, 40.2720 21.7628, 28.05.1985, HAUS (1) ; Kozani, 4.3 Km in dir. Florina , 1.3 Km N Kéla, 40.3552 21.797, 720 m alt., 24.9.1989, NMBE 524565 / 1; Kozani , 2 Km S Xirolimni, 40.2744 21.6465, 700 m alt., 14.5.2005, NMBE 523305 / 4; Xirolimni, 4.2 Km in dir. Kozani / Makedonia, 40.3164 21.7457, 700 m alt., 25.9.1989, NMBE 524569 / 2; Asprovalta , 40.7204 23.7057, 30.7.1984, NMBE 512667 / 1; Chalkidiki, ruins of Olynthos , 40.296 23.354, 50 m alt., 24.7.1990, NMBE 512652 / 2; Chalkidiki, Petralona at Eleohori , 40.344 23.162, 180 m alt., 28.8.1991, NMBE 512653 / 3; Chalkidiki, cave of Petralona , 40.374 23.165, 310 m alt., 23.7.1990, NMBE 512651 / 1; Chalkidiki, Sithonia, Porto Koufos , 39.962 23.933, 20, 8.9.1991, NMBE 512666 / 7; Chalkidiki : 5 km NW Krini, 40.401 23.111, 180 m alt., 28.8.1991, NMBE 512654 / 1; Chalkidiki, Eleohoria , 40.3434 23. 1613, 170 m alt., 5.5.1995, NMBE 524564 / 2; Asvestohori , 40.6466 23.0236, 400 m alt., 8.5.1997, NMBE 524568 / 2. Thessaly: SMF 74190/4 Thessaly, coll. Krüper; ditto, SMF 299393, coll. Kaltenbach; Castle of Nea Panteleimon S Katerini, 39.989 22.603, 200 m alt., 11.9.1991, NMBE 512655 / 5; SMF 280623/20, Oros Olimbos, Platamon, 39.9905 22.6244, coll. Schlickum ex Schütt ; SMF 280626/2 Canyon of Tyrnavos between Larissa and Elassona, 39.8091 22.2703, coll. Schlickum ex Schütt ; SMF 280625/2 Levadia , Lofistion between Levadia and Thiva, at the Kopais Lake , coll. Schlickum ex Schütt ; SMF 18552, Volos , 39.3621 22.9421, coll. O. Boettger ex Stussiner ; ditto, SMF 18547, coll. Kobelt ex Heldreich ; ditto, SMF 74211/3, coll. C.R. Boettger; ditto, SMF 78213/, coll. Nägele; ditto, SMF 74172/9, coll. Jetschin ex Stussiner; Volos, Platania on the Pilion, 39.1429 23.2750, 15.10.1986, LIEB (2) ; Volos, Mt. Pelio , 39.4384 23.0475, 1610 m alt., 1845, NMBE 22789 / 2; Volos, Chorefto on the Pilion , 39.4519 23.1200, 08.10.1986, LIEB (3) ; Volos, Makrinitsa on the Pilion , 39.4017 22.9877, 30.08.1986, LIEB (14) ; Pilion, rocks 4 km W Xania above Volos , 39.38 23.03, 990 m alt., 15.6.1994, NMBE 512656 / 2; Néa Ionia (= Volos ), W of harbour, 39.3492 22.9768, 30.4.1991, NMBE 524513 / 3; Kalambaka, Moni Megalo Meteoron , 39.7266 21.6257, 20.08.1982, HAUS (3) ; Trikala in direction to Larissa, rocks at crossroads Petrolon , 39.5650 21.9703, 17.06.1985, HAUS (3) ; Ampelia 1.5 km in direction to Kalambaka , 39.7082 21.5269, 20.06.1985, HAUS (1) ; Theopetra , 39.6737 21.6843 21.08.1982, HAUS (2) ; ditto, rocks close to the church, 21.08.1982, HAUS (6) ; Theopetra, Kuvelci, 6 Km SSE von Kalambaka, 39.6798 21.681, 260 m alt., 26.7.1975, NMBE 524525 / 6; Vlochos 1 km in direction to Palasmas 39.4956 22.0924, 17.06.1985, HAUS (2) ; ditto, 28.08.1982, HAUS (7) ; Vlochos 3.8 km in direction to Farkadon , aride rocky slope, 39.5487 22.0926, 17.06.1985, HAUS (3) ; Farkadon , 39.5933 22.0674, 17.06.1985, HAUS (5) ; SMF 74183/3 Lamia , coll. Krüper; Kastri, castle, 39.6246 22. 6905, 135 m alt., 12.5.2005, NMBE 523333 / 140; Astolofos, Larissa , 39.692 22. 7273, 230 m alt., 12.5.2005, NMBE 523331 / 22; Larissa, 1 Km N of Potamia, 39.6686 22. 7877, 135 m alt., 12.5.2005, NMBE 523312 / 1. Sterea Ellada: SMF 9707, SMF 18548/2, Hypati [Ipati], 38.8716 22.2413, (paralectotypes of pelasgica ), coll. Kobelt ex Thiesse; Ipati, waterfall, 38.871 22.244, 380 m alt., 16.5.1991, NMBE 524549 / 1; road 1.5 Km NE Larissa , 38.9673 22. 3874, 615 m alt., 10.4.1986, NMBE 524560 / 5; Dio Vouna, 3 Km in dir. Gorgopotamos , 38.8172 22. 3933, 360 m alt., 28.4.2003, NMBE 523329 / 2; Néo Monastiri, 6.5 Km in dir. Lamia, 39.1816 22. 2883, 140 m alt., 10.4.1988, NMBE 524558 / 1; Amfiklia , SE of village, chapel at canyon entrance, 38.6306 22.5963, 500 m alt., 10.4.1986, NMBE 524548 / 1; SMF 74168 Parnass, Grotto of Korykos , coll. K.L. Pfeiffer; Phthiotis, Pelasgia 1 km in direction to Glifa, 38.9453 22.8506, 17.10.1986, HAUS (2) ; Phthiotis, Pelasija 7.5 km in direction to Glifa, 38.9412 22.9142, 17.10.1986, HAUS (1) ; Mornou dam, 9 Km in dir. Lévka , 38.5342 22. 1634, 550 m alt., 13.4.1986, NMBE 524519 / 2; Agios Spirido , E of Glifada, 38.3679 22.1169, 30 m alt., 1.10.1989, NMBE 524553 / 8; Agios Spirido , E of village, 38.3681 22.1299, 60 m alt., 20.4.2003, NMBE 523326 / 3; Nafpaktos 28 km in direction to Galaxidi , 38.3637 22.1293, 25.08.1982, HAUS (3) ; Eratini , hill slope N of village, 38.3978 22.1931, 500 m alt., 14.4.1986, NMBE 524520 / 1; Lidoriki, 4 Km N crossroads to Eratini , 38.4384 22. 2387, 550 m alt., 13.4.1986, NMBE 524522 / 2; Glifada , hill slope W of village, 38.3883 22.0711, 15.4.1988, NMBE 524510 / 3; crossroads Lidoriki–Eratini–Amfissa , 38.4052 22.255, 640 m alt., 1.10.1989, NMBE 524537 / 1; Itea, 10 Km in dir. Galaxidi , 38.413 22.395, 15 m alt., 20.4.2003, NMBE 523301 / 6; ditto, 25.08.1982, HAUS (2) ; Itea, 6 Km in dir. Galaxidi , 38.4326 22.397, 20.4.2003, NMBE 523306 / 5; Gravia, 2.5 Km in dir. Amfissa, 38.6607 22. 4253, 540 m alt., 12.4.1986, NMBE 524550 / 1; Mariolada , S of village, 38.6517 22.4655, 400 m alt., 29.4.2003, NMBE 523308 / 4; SMF 74188/3, Delphi , 38.483 22.5, coll. Krüper; ditto, SMF 74167/3, coll. K.L. Pfeiffer; ditto, SMF 214869/4, coll. S. H. Jaeckel & Plate; ditto, SMF 311942/4, temple of Apollon, 23.10.1982, coll. Hohorst; ditto, 630 m alt., 16.5.1991, NMBE 524526 / 7; Antikira, 6.3 Km in dir. Desfina , 38.3889 22.5915, 500 m alt., 20.4.2003, NMBE 523324 / 3; Camping Apollon W Delphi , 38.4824 22. 4757, 480 m alt., 20.6.2003, NMBE 524420 / 1; canyon at the Castalian well, Delphi , 38.4831 22. 5055, 590 m alt., 21.6.2003, NMBE 512642 / 2; SMF 311943/4, monastery of Ossias Lukas , 38.3954 22.7467, 24.10.1982, coll. Hohorst; Itea, 38.428 22.456, 20 m alt., 22.6.1987, NMBE 512645 / 3; 5 km E Desfina , 38.4 22. 58, 520 m alt., 24.6.1987, NMBE 512646 / 2; road Eratini to Amfissa , 38.4 22. 27, 650 m alt., 25.6.1987, NMBE 512648 / 1; castle of Amfissa , 38.531 22.373, 230 m alt., 25.6.1987, NMBE 512647 / 1; SMF 311947/1, Eratini , 38.3606 22.2286, 23.10.1982, coll. Hohorst; Lidorikion 15.5 km in direction to Nafpaktos, 38.5185 22.1525, 11.06.1985, HAUS (2) ; Nafpaktos , castle, 38.3969 21.8241, 100 m alt., 12.6.2003, NMBE 524421 / 3; Kalesméno, Tavropos River , 38.9418 21.6832, 300 m alt., 17.5.1995, NMBE 524566 / 1; Paralia Sarandi, 4.3 Km in dir. Prodromos, 38.2578 22. 8826, 225 m alt., 15.4.1986, NMBE 524570 / 81; crossroads Distomo–Delfi–Livadia , 38.4627 22. 7161, 310 m alt., 14.4.1986, NMBE 524547 / 1; Kiriaki, 6 Km in dir. Distomo , 38.3625 22.7208, 100 m alt., 14.4.1986, NMBE 524532 / 1; Paralia Sarandi, 3 Km in dir. Prodromos , 38.2572 22. 8756, 220 m alt., 15.4.1986, NMBE 524530 / 1; Korini [Domvrena], SE of Thisbi, 38.2513 22. 9886, 170 m alt., 15.4.1986, NMBE 524541 / 3; mountain E Kalamaki , 38.2005 23.1261, 25 m alt., 1.5.2003, NMBE 523325 / 1; SMF 74194 Agoriani , 38.8851 22.1150; road cross Akréfnio, 5.5 Km SE crossroads at Kastro, 38.45 23. 2092, 120 m alt., 11.5.1997, NMBE 524562 / 2; Skoponéri , 38.5032 23.3549, 10 m alt., 11.5.1997, NMBE 524567 / 2; 2 Km E Aliartos , 38.4282 23. 5309, 270 m alt., 9.4.1988, NMBE 524531 / 2; Tithoréa , canyon and cave, 38.5702 22. 6602, 850 m alt., 11.4.1986, NMBE 524551 / 4; Krionéri , 38.3452 21.5969, 8.4.1986, NMBE 524512 / 4; ditto, steep rocks, 08.06.1985, HAUS (1) ; Mt. Varassova, SW slope, Krionéri , 38.3477 21.5973, 50 m alt., 1.10.1989, NMBE 524498 / 5. Attica: SMF 74187/12, SMF 74198 Attica, coll. Krüper; ditto, SMF 78210/2, coll. Ehrmann ex Schlüter; ditto, NMBE 22788 / 3; SMF 74175/4 Attica, Daphni, 38.0129 23.6358; SMF 18531 ruderal field E of Athens, coll. O. Reinhardt; SMF 74171/2 Ambelaki at Salamis , 37.9510 23.5280, coll. K.L. Pfeiffer; SMF 280622/8 Megara , 37.9952 23.3426, coll. Schlickum ex Schütt; Mégara, Kloster Agios Ierothéou / Atiki, 38.093 23.2677, 300 m alt., 7.7.1979, NMBE 524499 / 1; Mégara , W of city, 37.9962 23.324, 65 m alt., 7.7.1979, NMBE 524511 / 3; SMF 74165 Attica, Sunion , 37.6591 24.0147 coll. K.L. Pfeiffer; Aigosthena (= Porto Germeno), 38.1483 23.2321, 45 m alt., 16.5.1991, NMBE 524502 / 3; SMF 24476/4 Athens, 37.9839 23.7293 ex Raymond; SMF 18534/3 Athens , Hymettos , 37.9583 23.8196, coll. O. Boettger ex P. Hesse; SMF 18528/2, Athens , Lykabettos , 37.9826 23.7436, coll. Kobelt ex Anthes ; ditto, SMF 74169/2, coll. K.L. Pfeiffer; ditto, SMF 74199, K.L. Pfeiffer ex Kaltenbach ; ditto, SMF 299392/3, SMF 299555, coll. Kaltenbach; ditto, 250 m alt., 19.7.1979, NMBE 524523 / 6; ditto, 22.4.1988, NMBE 524503 / 2; Varkiza, Camping "Varkiza-Beach", 37.8226 23.817, 30 m alt., 20.7.1979, NMBE 524500 / 2; Vari, 3 Km in dir. Koropi , 37.8415 23.8215, 50 m alt., 20.7.1979, NMBE 524504 / 2; Keratéa, 1.5 Km in dir. Markopoulon, 37.8542 23.9345, 100 m alt., 20.7.1979, NMBE 524509 / 2; Lavrio, 4 Km in dir. Markopoulo , 37.7523 24.026, 125 m alt., 5.5.1991, NMBE 524501 / 1; SMF 311948/2, Athens, Philopoppos hill, 25.10.1982, coll. Hohorst; Mandra 2 km in direction to Thiva , 38.0851 23.4841, 16.10.1986, HAUS (1) ; rocks at a quarry in Kasa 2.5 km in direction to Erithres, 22.09.1986, HAUS (1) ; Kifissia , 38.08 23. 82, 260 m alt., 3.1864, NMBE 18463 / 4; ditto, 1864, NMBE 18469 / 3; Vouliagmeni , 37.8216 23.7754 07.01.1985, LIEB (1) .; SMF 18550/2 Isthmus of Korinth, 37.9257 23.0017, coll. O. Reinhardt. Peloponnese: SMF 74205 Peloponnese, coll. Krüper; SMF 78211 Korinth , 37.9375 22.9306, coll. Ehrmann ex Schlüter; Zemeno 5.5 km in direction to Kaliami, canyon, 14.10.1986, HAUS (1) ; Zemeno close to Xylokastron , rocks at the school, 38.0299 22.6059, 14.10.1986, HAUS (2) ; SMF 74186/3, Xylokastron , 38.0772 22.6326, coll. Krüper; Xilokastro between Pellini and Dendro, 38.0392 22.5393, 500 m alt., 11.7.1996, NMBE 512659 / 4; Kefalari 1 km in direction to Xylokastron , 37.9303 22.5316, 15.10.1986, HAUS (1) ; Stimfalia Limni , 37.87 22.47, 27.4.1996, NMBE 13584 / 3; Kaliani, 6 Km in dir. Kiato , Killini Mts. , 37.9236 22. 5262, 710 m alt., 8.7.1979, NMBE 524538 / 1; Kaliani, 10 Km in dir. Kiato , 37.9473 22. 5663, 780 m alt., 8.7.1979, NMBE 524544 / 8; Kefalari , mountain 2 Km E village, 37.9151 22. 5319, 730 m alt., 5.5.2005, NMBE 523332 / 55; Agios Vassilios, 8.5 Km in dir. Argos , 37.7779 22. 7339, 290 m alt., 18.7.1979, NMBE 524528 / 1; Akrokorinthos , old castle, 37.89 22.869, 400 m alt., 18.7.1979, NMBE 524543 / 7; Akrokorinthos , 37.8906 22. 8708, 520 m alt., 12.7.1996, NMBE 512660 / 4; Arhea Korinthos , 37.9059 22.8792, 80 m alt., 8.1973, NMBE 524536 / 1; Heraiou Lake, Heraion , 38.0349 22.8708, 60 m alt., 18.7.1979, NMBE 524539 / 1; lighthouse Akron Ireo, Heraion, Korinthia , 38.0285 22. 8499, 460 m alt., 12.7.1996 NMBE 512661 / 2; above Loutraki , 37.9801 22.9858, 90 m alt., 19.2.1981, NMBE 524542 / 1; Sikion close to Kiato , theatre, 38.0079 22.7444, 14.10.1986, HAUS (1) ; Nemea 1 km in direction to Kaliani , 37.8431 22.6392, 16.10.1986, HAUS (2) ; rocks SE of Nemea , 37.8110 22.6886, 16.10.1986, HAUS (1) ; Killini Mountains, Mesi Sinika Trikalon , 15.10.1986, HAUS (1) ; Neméa, 1 Km S / Korinthia , 37.8024 22.667, 370 m alt., 7.5.1991, NMBE 524527 / 1; Agios Vasilios , 37.7907 22.7974, 12.4.2003, NMBE 523303 / 1; ditto, 6.5.2005, NMBE 523322 / 2; SMF 74173/2, Kyllene [Killini], 37.9346 21.1449 coll. Jetschin; SMF 18549, Dimitsana, 37.5955 22.0395, coll. O. Reinhardt; SMF 74170, road Argos to Tripolis , on the pass, 37.5326 22.5818, coll. K.L. Pfeiffer; SMF 214868, Levidhi , 37.6820 22.2956, coll. S. H. Jaeckel; SMF 214895/2, Tripolitza [Tripolis], 37.5101 22.3726, coll. S. H. Jaeckel ex Martens; Paralio Astros, castle, 37.4183 22.7663, 50 m alt., 16.7.1979, NMBE 524557 / 1; ditto, 17.7.1996, NMBE 512665 / 1; Astros, 7 Km in dir. Argos , 37.4663 22.7421, 75 m alt., 16.7.1979, NMBE 524529 / 3; 3 Km S Astros , hill slope, 37.3841 22.7437, 20 m alt., 13.4.2003, NMBE 523323 / 4; Astros 3 km in dir. Leonidion , 37.3789 22.7455, 24.09.1986, HAUS (1) ; Vitina , 5 Km SW of village, 37.6515 22. 1685, 985 m alt., 17.4.1988, NMBE 524555 / 2; Levidi, 4.7 Km in dir. Kandila, 37.7214 22. 3276, 720 m alt., 7.5.1991, NMBE 524518 / 1; Perthori, 3.7 Km in dir. Moni Epano Hrépas, 37.5417 22.3347, 1000 m alt., 8.5.1991, NMBE 524534 / 2; Manthiréa, 4 Km in dir. Sparti , 37.3813 22. 4034, 860 m alt., 5.8.1976, NMBE 524540 / 1; Leonidion, Wof village , 37.1684 22. 8427, 145 m alt., 19.2.1981, NMBE 524554 / 10; ditto, 16.7.1979, NMBE 524545 / 11; Steno 0.5 km in direction to Tripolis, 37.4953 22.4554, 07.10.1986, HAUS (4) ; Rizes 1 km in direction to Ag. Petros, under stones at the crossroads to Astros, 37.4405 22.4880, 06.10.1986, HAUS (1) ; Stavrodromio, 1 Km in dir. Arhea Olympia , 37.6989 21. 9392, 630 m alt., 2.8.1976, NMBE 524516 / 1; Tropea, 6 Km N in dir. Dafni , 37.7594 21. 9578, 550 m alt., 17.4.2003, NMBE 523309 / 3; Dimitsana , 4 Km N of village, 37.6254 22.0606, 990 m alt., 16.4.2003, NMBE 523320 / 2; Kalonéri, 3.5 Km in dir. Arhea Olympia, 37.6719 22.0617, 1100 m alt., 16.4.2003, NMBE 523327 / 1; Ipsous (= Stemnitza ), 37.5515 22.0706, 1080 m alt., 16.4.2003, NMBE 523321 / 2; Pirgaki , 38.1648 22.0822, 700 m alt., 19.4.2003, NMBE 523302 / 4; Klitoria, 2 Km S in dir. Tripoli , 37.8808 22.1355, 500 m alt., 9.7.1979, NMBE 524521 / 1; Agios Vassilios, 2.7 Km in dir. Tripotama, 37.8629 21.9235, 600 m alt., 17.4.2003, NMBE 523307 / 3; Petsaki , canyon 1 Km N of village, 38.1009 22.0331, 905 m alt., 19.4.2003, NMBE 523319 / 1; Ptéri , 3 Km S, 38.1418 22.071, 1150 m alt., 15.4.1988, NMBE 524514 / 1. Argolis: SMF 18544 Argolis, Nauplion [Nafplio], 37.5673 22.8015, coll. O. Reinhardt; ditto, 25.3.1987, NMBE 512673 / 2; Nafplion , northern boundary, 08.10.1986, HAUS (2) ; Nafplion, Palamidi , 37.5618 22. 8042, 180 m alt., 12.7.1985, NMBE 524419 / 4; Nafplio , below castle, 37.5634 22.8013, 50 m alt., 18.7.1979, NMBE 524552 / 21; SMF 78212/6, castle of Mykene , 37.73 22. 76, 250 m alt., coll. Ehrmann ex Schlüter; ditto, SMF 299391/4, coll. C. Bosch ex Schlüter; ditto, SMF 311946/2, 26.10.1982, coll. Hohorst; ditto, NMBE 512658 / 1, NMBE 18467 / 2; Mykene , valley below castle, 37.7221 22. 7538, 150 m alt., 20.4.1988, NMBE 524563 / 27; SMF 74166/3, Epidavros , 37.5958 23.0768, coll. K.L. Pfeiffer; ditto, SMF 311945/1, theatre of Epidavros, 27.10.1982, coll. Hohorst; Lygourio 3 km in direction to Nea Epidavros, 37.6319 23.0536, 13.10.1986, HAUS (1) ; Didima , rocks righ of the doline, 37.4728 23. 1648, 540 m alt., 13.7.1996, NMBE 512662 / 1; Didima , 37.4639 23.1725, 10.10.1986, HAUS (2) ; Argos , rocks above the church, 37.6351 22.7288, 23.09.1986, HAUS (1) ; Argos, castle Larissa , 37.6383 22. 7154, 250 m alt., 16.7.1996, NMBE 512664 / 5; ditto, 18.7.1979, NMBE 524559 / 5; rocky slopes N of Argos , 22.09.1986, HAUS (1) ; Agrilitsa W of Argos, 37.6246 22.5964, 23.10.1986, HAUS (1) ; Argos, Aspis hill, 23.09.1986, HAUS (1) ; bay 5 km W Iria , 37.512 22.9817, 16.7.1996, NMBE 512663 / 1; Agiorgitika, 12.5 Km in dir. Argos , 37.5385 22.5741, 800 m alt., 5.4.1986, NMBE 524561 / 3; Ahladokambos, 7 Km in dir. Argos , 37.5168 22. 6154, 650 m alt., 20.4.1988, NMBE 524533 / 1; Kenchreai , 37.8841 22.9859, 17 m alt., 18.7.1979, NMBE 524625 / 22; Vavari , 37.537 22.9211, 25 m alt., 21.5.2011, NMBE 523328 / 1, NMBE 523310 / 7; Kandia , 37.5288 22.9554, 17 m alt., 21.5.2011, NMBE 523311 / 3; Kranidi 1.5 km in direction to Ermioni, 37.3742 23.1762, 11.10.1986, HAUS (1) ; eastern boundary of Vivarion , 10.10.1986, HAUS (1) ; quarry opposite to Kilada , 37.4133 23.1253, 11.10.1986, HAUS (1) ; Ermioni 1.2 km in direction to Kranidi , 37.3846 23.2329, 11.10.1986, HAUS (3) ; slope at the road Ag. Triada in direction to Arachneo, 37.6508 22.9144, 08.10.1986, HAUS (2) ; Amarianos 2 km in direction to Arachneo , 37.6470 22.9033, 08.10.1986, HAUS (1) ; SMF 311944/1, road Nafplio to Tripolis at the crossroads to Achladokambos , 37.5250 22.5901, 25.06.1985, coll. Hohorst; plateau above Gerolimenas, 36.486 22.387, 100 m alt., 7.7.1987, NMBE 512649 / 4; castle of Monemvasia , 36.687 23.052, 100 m alt., 10.7.1987, NMBE 512650 / 10; ditto, 30.09.1986, HAUS (3) .; Monemvassia, 2 Km in dir. Nomia , 36.671 23.0306, 17.2.1981, NMBE 524506 / 1; Geroliménas , 36.4842 22.3993, 25 m alt., 13.7.1979, NMBE 524508 / 1; Skala, 3 Km in dir. Sparti , 36.8594 22.6058, 100 m alt., 14.7.1979, NMBE 524505 / 2; crossroads Geraki–Sparti, Skála–Kosmás , 37.0607 22.7071, 800 m alt., 18.2.1981, NMBE 524524 / 15; Dermatianika, Mt. Kabos , 36.5451 23.0533, 70 m alt., 17.2.1981, NMBE 524507 /1.

Greek Islands: Agios Eustratios: Agios Eustratios, 39.5368 24.9897, 21.05.1986, LIEB (1). Donousa: Donousa, 37.1005 25.7951, 02.05.1984, LIEB (9). Skopelos: SMF 74203/1, SMF 74179, Skopelos, 39.1251 23.6799, coll. Krüper. Skiathos: Megalos Aselinos, 39.1720 23.4226, 01.06.1979, LIEB (45). Skyros: Molos, 38.9147 24.5655, 01.10.1982, LIEB (4). Evvia: SMF 74197 Evvia, coll. Krüper; ditto, SMF 18536/3, ex Godet; ditto, SMF 74174/2, coll. Jetschin ex Schneider; ditto, SMF 74204, coll. Krüper; Halkida (= Chalkis), 38.47 23.62, 1884, NMBE 18466/2; 2 km S Pilio, 2 km in dir. Vlahia, 38.7579 23. 6185, 170 m alt., 18.6.1994, NMBE 512657/1; Southern slope of Ochi Mts., 5 km E of Bouros, 37.9512 24.5005, 35 m alt., 31.09.2000, NMBE 512643/1; Mt. Olimbos, NE slope, small canyon E of Mistros, 38.5183 23.8591, 320 – 600 m alt., 04.10.2000, NMBE 512669/2 (alk.), NMBE 512644/7; Mantoudi, 38.7942 23.477, 20 m alt., 1.5.1991, NMBE 524515/3; Néa Artaki, 1–2 km N of village, 38.5404 23.6344, 9.4.1988, NMBE 524556/1; Alivéri, 38.4022 24.0138, 350 m alt., 16.4.1986, NMBE 524517/3; Vrissi, Mavrovouni Mts. cave niches, 38.5549 24.0268, 450 m alt., 16.4.1986, NMBE 524546/1. Samothraki: Chora, 40.4715 25.4751, 10 m alt., 11.5.2008, NMBE 523304/7. Keos: Otzias, 1 Km E, 37.6774 24. 3625, 180 m alt., 6.5.1991, NMBE 524535/1. Naxos: SMF 74185/3 Naxos, 37.0755 25.5207, coll. Krüper; ditto, SMF 18545; ditto, Naxos, 04.09.1984, LIEB (6). Syros: SMF 74189/4 Syros, 37.4234 24.9160, coll. Krüper. Ägina: Ägina, 37.7408 23.5014, 03.06.1083, LIEB (1). Poros: SMF 74180/5 Poros, 37.5088 23.4586; ditto, SMF 74184/7, coll. Krüper. Paros: Paros, eastern coast at Piso Livadia, 37.036 25.2598, Sept. 1986, NMBE 512668/3, ex Kleinhenz; ditto, 3.1994, NMBE 505542/3. Limnos: Poliochni, 39.9093 25.1994, 31.05.1986, LIEB (1). Thira: Thira, 36.3931 25.4615, 19.05.1991, LIEB (11). Milos: Milos, 36.6914 24.3935, 15.06.1991, LIEB (1).

Turkey: SMF 74212/3, Adrianopel [Edirne], 41.6818 26.5622, coll. C.R. Boettger ; ditto, SMF 78209/19, coll. Nägele ex Braun; Balikesir Ili: Gulf of Edremit, AltIn Oluk , olive plantation, 39.5823 26.7393, 22.09.2001, LIEB (2) ; AltIn Oluk, limestone rocks at the Şahin Dere , 39.5844 26.7565, 22.09.2001, LIEB (3) . Izmir Ili: Pergamon , 39.1321 27.1843, 30.4.1992, leg. Tannert, ex coll. Groh, NMBE 512622 / 4; Bergama, Asklepion , 29.09.2001, LIEB (30).

Unknown localities: SMF 74182 Seidikeri.

Unspecific localities: SMF 18546 Greece, coll. O. Reinhardt

(*) Specimens from these localities show reticulate pattern on the shell surface and are similar to H. nucula .

Diagnosis: shell small, thin, spherical, protoconch small, teleoconch smooth with coarse axial riblets, aperture always white; flagellum as long as epiphallus+penis, long diverticulum exceeding bursa copulatrix in length.

Description: shell of small size, thin, spherical, spire elevated conical; basic shell colour greyish white with chestnut brown spirals, usually spiral No. 2 fused with No. 3 or diffuse, often with all five spirals separate; protoconch small, its diameter ranging from 3–4 mm, smooth, white whorls; teleoconch of 3–4 smooth whorls, sculptured by coarse but densely arranged axial riblets; last whorl slightly descending towards the periphery; aperture rounded, apertural rim narrow, labial callus weak, always white, columellar triangle small; umbilicus closed in adult shells.

Genital organs ( Figs 231–232 ): penis club-shaped, epiphallus considerably longer than penis; flagellum as long as epiphallus+penis; mrp attaching in a subterminal position on the epiphallus; internally, pp1 smaller than pp2, penial chamber long, filled with numerous minute ring-like folds, both papillae conical with a central perforation; atrial stimulator a rounded knob; female system with a very short vaginal tube, dart sac strongly developed, glandulae mucosae with a thick and short basic stem and a few short tubules, glandulae mucosae shorter than dart sac; pedunculus stem long, with a long diverticulum branching off in the upper third of pedunculus and exceeding bursa copulatrix in length, bursa copulatrix stem shorter than pedunculus stem, vesicle rounded and bent downwards.

Distribution ( Fig. 233 ): This species is widespread in Greece, and also reaches the northern slopes of the Rhodope Mountains in Bulgaria; however, its northern distribution boundaries are not known because of probable confusions with species living syntopic like H. philibinensis and H. pomacella . Hudec & Vasatko (1973: 30, Abb. 28) illustrate the genital organs of a specimen collected in the valley of the Marica river close to Biser at Harmanli (41.8619 25.9797), which match the major characteristics of specimens from Greece. The elongate epiphallus is clearly visible, with a relatively short flagellum, and the very long and well developed diverticulum. It is also recorded from the west Mediterranean Turkish coast in the area from Bergama southwards (Korábek et al., submitted).

Remarks: This species can best be differentiated from its next relative H. nucula by the absence of any granulation on its teleoconch. After study of hundreds of shells from Greece, only a few exceptions could be found, where some fine subsutural furrows were present. On the other hand, many shells of H. nucula from Turkey need very close examination for the granulation, which is sometimes only visible under high magnification. In most shells of H. nucula , the granulation pattern is restricted to a small subsutural area on the upper whorls, but sometimes it can cover large parts of the teleoconch. The character states of the genital organs are very similar in both species; the stems of the glandulae mucosa and the vaginal tube seem to be somewhat longer in H. nucula than in H. figulina , and probably the branching point of the diverticulum is different in both species. However, it is not clear whether these slight differences are good character states to be used for species discrimination, or if they simply reflect individual variation.

In shell characters, H. figulina differs from both, H. philibinensis and H. pomacella by its very small protoconch. Helix philibinensis has a somewhat larger shell, usually a cream to brown aperture, and all colour spiral bands well separated. Shells of H. pomacella are considerably smaller, and much more globular. Problems in identification may arise in southern Bulgaria, while in Central Greece this species is unmistakable.

SMF 18540/3 Cerigotto, coll. O. Boettger ex Krüper; SMF 74181/3 Cerigotto, coll. Krüper: The specimens of this lot are determined as figulina , but the shells are heavy, broader and have a protoconch> 5 mm. The shells are subfossil/fossil, their identity is not clear yet. SMF 74185/3 from Naxos: These specimens are very small and globose and resemble pomacella .

Figures

Figs 227–230. Helix (Pelasga) figulina ROSSMÄSSLER, 1839. Fig. 227 holotype figulina: SMF 9845, Griechenland, D = 22.1; Fig. 228 lectotype pelasgica: SMF 9706, Hypati in der Phthiotis, D = 31.2; Fig. 229 NMBE 512669, Mistros, D = 29.2; Fig. 230 ditto, D = 28.4. — All photos Neubert, original size.

Figs 231–232. Helix (Pelasga) figulina ROSSMÄSSLER, 1839. Genital organs. Fig. 231 NMBE 512669, Mistros, situs 43.1; Fig. 232, ditto, situs 45.7.

Fig. 233. Helix (Pelasga) figulina ROSSMÄSSLER, 1839. Distribution.

Figs 1–9. Helix (Helix) pomatia LINNAEUS, 1758. Fig. 1 compacta, syntype SMF 9573, D = 50.07, Hungary "Gestrüppe der Berglehnen"; Fig. 2 pulskyana, syntype SMF 9525/2a, Hungary "Inseln der Donau", D = 36.6; Fig. 3 hajnaldiana, syntype SMF 9658, Hungary "Budapest, im botanischen Garten", D = 34.1; Fig. 4 solitaria, syntype SMF 74708/2a, D = 31.1; Fig. 5 sabulosa, syntype SMF 9524, D = 40.5, Fig. 6 expansilabris, holotype SMF 9538, D = 45.5; Fig. 7 luteola, holotype SMF 9527, D = 34.6; Fig. 8 pannonica, lectotype SMF 9535, D = 40.5, Fig. 9 lednicensis, syntype SMF 9534/2a, D = 38.3. — All photos Neubert, original size.

Figs 10–19. Helix (Helix) pomatia LINNAEUS, 1758. Fig. 10 claudiensis, holotype SMF 9543, Klausenburg, D = 39.5; Fig. 11 dobrudschae, holotype SMF 9541, Rumania, Dobrudscha, Comana Vlasca, D = 46.2; Fig. 12 costellata, holotype SMF 9631, Markt in Bukarest, D = 39.7; Fig. 13 elsae, lectotype SMF 9529, nahe Bukarest, D = 34.9; Fig. 14 transsylvanica, holotype SMF 9634, Markt in Bukarest, D = 40.1; Fig. 15 christinae, lectotype SMF 9542, Runku über Bukarest, D = 44.5; Fig. 16 kapellae, holotype SMF 9544, Kapella Gebirge, D = 44.2; Fig. 17 serbica, holotype SMF 9540, Maidanek, D = 53.9; Fig. 18 thessalica, lectotype SMF 9568, Makrinitza bei Volos, D = 45.4; Fig. 19 rhodopensis, syntype SMF 9599/2a, Philippoppel, D = 41.0. — All photos Neubert, original size.