Cochlostoma (T.) macei ( Bourguignat, 1870 )

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 127

Cochlostoma (T.) macei ( Bourguignat, 1870)

Figs 105 (white dots), 111

Pomatias macei Bourguignat 1870: 56–57 .

Types

Not seen.

Other specimens

FRANCE • 1- Gourdon ; 43.7305° N, 6.959° E; 1976; RMNH leg.; EZ0399 • 2- Gorge du Trevans ; 43.9471° N, 6.2267° E; 2014; Zallot leg.; EZ1178 .

Type locality

FRANCE • Drôme, Grasse, Saint Vallier.

Description

SHELL. Closely spaced riblets on last part of protoconh. Teleoconch spotless with moderately strong ribs. Relatively strong lip with quite abruptly inwardly curved columellar lobe covering umbilicus.

MEASUREMENTS. 3 ♀♀: whorls= 8.3–9.1, H = 9.4–10.4 mm, H/W=2.79–3.1, roundness =0.1–0.14, ribs incl.= 57–59°, apert. incl.= 17–19°, ribs/mm 1 st wh.= 5–13, ribs/mm 4 th wh.=7–15.

FEMALE GENITAL ORGANS. 2 specimens analysed. Slightly posterior connection of pedunculus to bursa copulatrix. Short, roughly oval seminal receptacle with undefined basis. 2–4 loops clustered beside apex of seminal receptacle. Junction of uterus gland far from connection of pedunculus of bursa copulatrix with distal oviduct.

Remarks

Our sample closest to the type locality originates from Gourdon, ca 10 km east of Saint Vallier. The other sample, from the Gorge du Trevans, is located 50 km northeast of the type locality. Only of the latter we were able to analyse the genitals.

The shell morphology and the female genitalia suggest that this species belongs to Turritus . Only H3 was sequenced and the results support this conclusion. Based on some similarity of the female genital morphology with that of C. (T.) sardoum , it could belong to the same subclade within clade B. This would be interesting in a paleogographic perspective because it would be the ‘remnant’ on the European plate of the closest relative to the sardoum , agriotes / affine , and atlanticum taxa (see Discussion), which originated on the southern border of the European plates before the geological eras which led to the current shape of the western Mediterranean Sea and the lands that sourround it.

Figures

Fig. 105. Distribution of samples with undetermined position in the phylogenetic trees: white= Cochlostoma (T.) macei (Bourguignat, 1870); purple =C. (T.) sturanii (Wagner, 1897); orange=C. (T.) kleciaki (Braun, 1887); yellow=C. (T.) arnautorum (Wagner, 1906); red =C. (T.) atlanticum (Bourguignat, 1868); green =C. (T.) euboicum (Westerlund, 1885); blue =C. (T.) martensianum (Möllendorff, 1873); magenta= C. (T.) simrothi (Caziot, 1908); cyan =C. (T.) fontqueri Haas, 1924.