Agra granodeoro Erwin

Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), Zootaxa 119, pp. 1-68 : 45-46

Agra granodeoro Erwin , new species

( Figs. 19a, 19 b, 19c)

Holotype. Male, COSTA RICA: Cartago, Turrialba, Chirripo, Grano de Oro, 1120 m, 09° 53 'N 83°38'W, September, (P. Campos)( INBio: CRI 000­935857).

Diagnosis. This is the smallest member of the palmata species­group in Costa Rica and the only species with a narrow, tapered prothorax, black knees and tarsi.

Description. Color and luster: Piceous. Legs rufotestaceous with black knees and tarsi. Antennomeres 1­2 and apex of 3­10 infuscated. Female unknown. Form: Head ( Fig. 19a ) behind eyes moderately long, sides tapered to abruptly obtuse angles. Pronotum widest at base, markedly narrowed just posterior to apical collar; disc sparsely punctate. Elytron with apex ( Fig. 19 b) markedly oblique, finely dentate suturally, sutural margin sinuate. Aedeagus ( Fig. 19 c) with two ostial membranes separated by a wide sclerotized bridge, endophallus without vestiture or armature; apex of phallus diamond­shaped. Size: moderately large, 18.1 mm in length, 4.4mm in width.

Other specimens examined. The male Holotype is the only specimen I saw.

Specific epithet. The specific epithet, granodeoro , is used as a noun in apposition, and refers in Spanish to a "golden seed," in reference to the brassy glint of the elytra and to the type locality.

Notes. The Holotype is from the Cordillera Volcánica Central Conservation Area. An illustration of palmata­group stylomere 2 is on the Agra atriperna web page.

Figures

FIGURE 19. A. granodeoro, male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0.