Calosoma (Carabomimus) sylvestre ( Lassalle, 2009 )

Häckel, Martin, Heinz, Walter & Sehnal, Rostislav, 2011, Revision of the Mexican subgenus Carabomimus Kolbe, 1895 (Coleoptera: Carabidae: Calosoma Weber, 1801): C. depressicolle, C. flohri and C. digueti species groups, Zootaxa 2908, pp. 45-63 : 51-52

3. Calosoma (Carabomimus) sylvestre ( Lassalle, 2009) , stat. n.

(Plate 2: Fig. 9)

Calosoma depressicolle Chaudoir, 1869 , loc. Río Frío ( Breuning, 1928, p. 375., coll. Mus. Berol.). Carabominus depressicollis sylvestris Lassalle, 2009, p. 3, Type: Mexique, México, Rio Frio, route México-Puebla, 2950 m, in coll. Lassalle.

Holotype: 13, Mexique, México, Rio Frio, 2950 m, route México – Puebla, VII.2007, in coll. Lassalle. Paratypes: 303, 30Ƥ, the same data as holotype, except 2700–3050 m, in coll. Van den Berghe and Lassalle.

Material studied. 89 specimens—13, “ MÉXICO México / Río Frío [de Juarez] / VI-1991, ex coll. W. N. Johnson [white label, printed] (Plate 2: Fig. 9, MHCH); 1 Ƥ, “ Mexique. Foret de Rio Frio / env. près statt trés. Puebla, 3000 m / coll. Mateu [white label, printed] ( WHCS); 1Ƥ, same data as holotype except VIII – 1978, ex coll. E. van den Berghe ( MHCH); 33, 5Ƥ, same data as holotype except Llano Grande/ Río Frío, ~ 2800 m, VII-1990, lgt. W. Heinz; 53, 8Ƥ, same data as holotype except Llano Grande/ Río Frío, ~ 2800 m, VII-1991, lgt. W. Heinz; 23, 6Ƥ, same data as holotype except Llano Grande/ Río Frío, ~ 2800 m, VIII-2000, lgt. W. Heinz; 263, 26Ƥ, same data as holotype except Llano Grande/ Río Frío, ~ 2800 m, VIII-2003, lgt. W. Heinz (Fig. 15, MHCH); 43, 1Ƥ, 30 km W of Texmelucan, Río Frío ( MHCH, RSCU, VBCP, WHCS).

Description. A small Calosoma but relatively large for the subgenus Carabomimus . Habitus similar to that of C. depressicolle Chaudoir , but specimens larger and relatively wider. Length including mandibles 20.2–22.5 mm (males) and 21.8–23.6 mm (females), width 7.5–8.4 mm (males) and 8.5-9.0 mm (females). Dorsum uniformly black, appendages and venter dark brownish black to black. Head normal, not hypertrophied, subquadrate, eyes remarkably prominent. Labrum bilobed and distinctly divergent anteriorly, with two lateral setae, its apical margin strongly emarginate. Clypeus with lateral margins strongly convergent towards apex, apical margin moderately emarginate. Frontal furrows indistinct, barely discernible. Dorsal surface of head sparsely but coarsely punctate, especially from clypeus to posterior margin of eyes. Mandibles stout, either smooth or finely strigose and punctate. Antennae long, reaching beyond humeri and to body midlength; third segment moderately compressed at base, second segment and base of fourth segment show virtually no compression. Pronotum narrowed posteriorly, more than one and half times wider than long, widest anterior to midlength; sides anteriorly arcuate and posteriorly rather straight; hind angles narrowly rounded and extending beyond basal line; lateral margin thin, slightly wider at base; entire surface smooth. Elytra with more or less regular, slightly elevated intervals, primaries with distinct rows of large but rather irregularly set foveae, secondaries and tertiaries interrupted and sometimes lost in coarse granulation. Aedeagus (Plate 2: Fig. 9) differs from that of C. (Carabomimus) depressicolle Chaudoir, 1869 by its more robust form; preapical widening is more pronounced relative to apex proper. Ecology. Specimens of the new species were collected together with C. (Carabomimus) costipenne Chaudoir, 1869 , at elevations of approximately 2500–2900 m, on partially forested northern slopes in the northern part of the volcanic Popocatepetl massif, around a pass between the peaks El Mirador in the north and Iztaccihuatl in the south.

Characters that justify the ranking of this taxon to the species level are the structure of the aedeagus and the overall habitus, which differentiate this species from others included in the depressicolle group.