Empis (Coptophlebia) atratata, Daugeron & Grootaert, 2003

Daugeron, Christophe & Grootaert, Patrick, 2005, Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae), Zoological Journal of the Linnean Society 145 (3), pp. 339-391 : 360

EMPIS (COPTOPHLEBIA) ATRATATA CLADE

Monophyly: The monophyly of this clade is supported by two characters: epandrium with a dorsal double brush of spine-like bristles [character 30(1)], and epandrium with at least 1 very strong, long bristle at tip [32 (1)].

Distribution ( Fig. 16 ): This group of three species is typically South-east Asian, recorded from Myanmar and Thailand.

Remarks: Although E. (C.) velutinella Frey is not included in the phylogenetic analysis and not reviewed taxonomically, a preliminary morphological study allows inclusion of this species in the E. (C.) atratata clade as well as in the key given below. A fourth species from China (Yunnan) with 2 very strong, long bristles at the tip of the epandrium seems to also belong to this clade. It is known from 1 male and 2 females (KBIN–IRScNB) but not described here because of the poor condition of the specimens.

Figures

Figure 16. Distribution of the Empis (Coptophlebia) atratata clade: E. (C.) atratata Daugeron & Grootaert = 2; E. (C.) kosametensis Daugeron & Grootaert = 2; E. (C.) velutinella Frey = 1.