Boreantrops subfoveolatus, Kits, Joel H. & Marshall, Stephen A., 2015

Kits, Joel H. & Marshall, Stephen A., 2015, A revision of Boreantrops Kits & Marshall (Diptera: Sphaeroceridae: Archiborborinae), Zootaxa 3915 (3), pp. 301-355 : 330-332

Boreantrops subfoveolatus sp. nov.

( Figs. 61, 62 , 75, 118)

Diagnosis. Separable from other members of the mexicanus subgroup by the following combination of characters: Prementum orange. Leg joints narrowly orange. Mid tibia with a row of anterodorsal bristles. Mesoscutum shallowly sculptured. Tarsi yellow.

Description. Head orange, maxillary palp yellow. Mostly covered with microtomentum, frons with microtomentum medially reaching from postocellar bristles anteriorly through ocellar triangle to anterior margin of frons, microtomentum on orbital plates reaching bases of interfrontal setae, face with patch of microtomentum below lunule and antenna. Ocellar bristles at level of anterior margin of median ocellus. Subvibrissal bristle and anterior genal bristles about 0.5X length of vibrissa.

Thorax black, mostly covered with microtomentum. Proepisternum shiny, anepisternum with microtomentum along dorsal third and posterior third, katepisternum with a shiny spot behind fore coxa, meron shiny. Mesoscutum shallowly sculptured. Halter white, brownish below knob.

Legs black, trochanters and joints reddish brown, tarsi yellow. Mid tibia with row of anterodorsal, 1 anteroventral, 1 posteroventral, and 5 subapical bristles. Hind tibia with 1 thin anteroventral, 1 ventroapical bristle.

Wing brown, crossveins r-m and dm-cu paler.

Abdominal syntergite 1+2 heavily sclerotized with a semicircular weakly sclerotized patch anteriorly, black, covered with microtomentum. Tergites 3–5 weakly sclerotized, tergites 3–4 with heavily sclerotized bars. Sternites weakly sclerotized.

Male postabdomen: Sternite 5 parallel-sided near base, wider distally, slightly longer than wide, posterior margin with a deep notch and interrupted row of setae, anterior apodeme broad, as long as or slightly longer than external portion, without keel ( Fig. 62 ). Surstylus paddle-shaped. Hypandrial arms with ventral tab-like structures. Pregonite small, discrete. Postgonite narrowed above lobes, lobes widely separated, anterior lobe broad, rounded, posterior lobe pointed. Basiphallus with long epiphallus, short, broad, transparent preepiphallus. Distiphallus with spinose dorsal tube strongly curved near base, distal portion flattened, extending well beyond ventral portion of distiphallus ( Fig. 61 ).

Female postabdomen: Tergites 6–7 with anterior and lateral margins heavily sclerotized, center and posterior margin not sclerotized. Tergite 8 covered with microtomentum, except anterior margin. Epiproct without anterior arms, mostly covered with microtomentum except anterior corners, with 2 setae. Cerci broad. Sternites 6–7 with margins heavily sclerotized, center not sclerotized. Sclerites of sternite 8 broadest posteriorly, posterior half covered with microtomentum. Hypoproct lozenge-shaped, covered with microtomentum. Spermathecae about 3.5X longer than wide, annulated, apex not invaginated, sclerotized duct about as long as bulb (Fig. 75).

Type material. Holotype ♂: PANAMA: Chiriquí: Cerro Punta, 5 km ESE, 2600 m, dung traps, 23–28 May 1977, S. Peck, debu01040227 ( DEBU). Paratypes: same data as holotype (1 ♂, 1 ♀, DEBU).

Distribution. Only known from the type locality in Panama ( Fig. 118 ).

Etymology. The species name refers to the weakly sculptured mesoscutum, which has a shallowly pitted (foveolate) appearance. The name is a declinable adjective.

Comments. The female paratype contained about 15 eggs, each about half as long as the abdomen.

Figures

FIGURES 61 – 66. Boreantrops male terminalia. B. subfoveolatus: 61) phallus, postgonite, and phallapodeme, lateral view, 62) sternite 5; B. suchixtepecensis: 63) phallus, postgonite, and phallapodeme, lateral view, 64) sternite 5; B. zacapa: 65) phallus, postgonite, and phallapodeme, lateral view, 66) sternite 5.

FIGURES 115 – 120. Boreantrops distribution maps. 115) B. masneri (diamonds), B. calceatus (circle); 116) B. oaxacensis; 117) B. peruvianus; 118) B. subfoveolatus; 119) B. suchixtepecensis; 120) B. zacapa. Elevation is indicated by grey shading at 500 m intervals.