Galeopsomyia myrciae Hansson, 2023

Hansson, Christer & Hanson, Paul E., 2023, HYMENOPTERA: CHALCIDOIDEA), 5 The genus Galeopsomyia Girault by Christer Hansson * & Paul E. Hanson **, Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 3 (1), pp. 1-743 : 400-401

Galeopsomyia myrciae Hansson sp.nov.

( Figs 9, 522, 523, 844, 986)

Diagnosis (female). Antennal clava with a distinct constriction between C1 and C2 ( Fig. 844); genal carina present; mesoscutellum ( Fig. 522) with strong reticulation on median part and with a weak median groove, lateral parts with strong reticulation, meshes elongate, submedian grooves distinct, ±straight and parallel, subdivided by transverse carinae; dorsellum with median carina; propodeum ( Fig. 522) with strong reticulation, callus with six setae; petiole very short, just a narrow band; gaster ( Fig. 522) 2.7× as long as wide, medio-basal Gt1 with a blunt edge shaped like an inverted U (as in Fig. 24), tergites with strong reticulation.

Female holotype: length of body 2.6mm (paratypes 2.0– 2.9mm).

Scape yellowish-brown, pedicel and flagellum brown. Face below level of toruli golden-green, clypeus dark brown, frons metallic bluish-green, antennal scrobes black; vertex metallic bluish-green. Mesoscutum and mesoscutellum golden-green, propodeum golden-red. Legs with fore and hind coxae golden-green, mid coxa dark brown; trochanters yellowish-brown; fore and mid femora yellowish-brown, hind femur with basal half dark brown and apical half yellowish-brown; tibiae yellowish-brown; T1–3 yellowish-brown, T4 pale brown. Petiole black. Gaster with Gt1 golden-green, Gt2 metallic bluish-green, Gt3-6 golden-purple medially and metallic bluish-green laterally, Gt7 black with metallic tinges; gonoplac black.

Antenna with a distinct constriction between C1 and C2. Face below level of toruli with weak reticulation, clypeus smooth, frons rugose, antennal scrobes with weak reticulation; with genal carina. Vertex with strong reticulation outside ocellar triangle, with wrinkled sculpture inside ocellar triangle; with a carina between posterior ocelli.

Mesoscutum with strong reticulation, meshes isodiametric on sidelobes, elongate on midlobe, midlobe rugose close to notauli; midlobe with a complete median groove, with 12 scattered adnotaular setae. Mesoscutellum with strong reticulation on median part and with a weak median groove, lateral parts with strong reticulation, meshes elongate; submedian grooves distinct, ±straight and parallel subdivided by transverse carinae; with 12 setae scattered over lateral parts. Dorsellum with strong reticulation, with median carina. Propodeum with strong reticulation; callus with six setae. Fore and hind coxae with strong reticulation, mid coxa with weak reticulation. Fore wing with six setae on dorsal surface of submarginal vein; speculum closed; costal setal row unbroken.

Petiole very short, just a narrow band. Gaster elongate; medio-basal Gt 1 with a blunt edge shaped like an inverted U; tergites with strong reticulation.

Relative measurements: head length, dorsal view 26.5; head length, frontal view 42; POL 10; OOL 6.5; lateral ocellus diameter 6; head width 58; mouth width 17.5; malar space 13.5; eye length 26; scape length 20; scape width 5; pedicel+flagellum length 60; pedicel length 7.5; pedicel width, dorsal view 4.5; F1 length 10; F1 width 5; F2 length 10; F2 width 5.5; F3 length 10; F3 width 5.5; clava length 22; clava width 6; C3 length 7; spicule length 2; mesosoma length 73; mesosoma width 52; midlobe of mesoscutum length 26; mesoscutellum length 30; mesoscutellum width 25; median part of mesoscutellum width (measured medially) 10; median part of mesoscutellum, width in anterior part11; median part of mesoscutellum, width in posterior part 10; lateral part of mesoscutellum, width (measured medially) 5.5; dorsellum length 5; propodeum length 11; costal cell length 38; costal cell width (measured at widest part) 2.5; marginal vein length 40; stigmal vein length 11.5; gaster length 115; gaster width 43; Gt 2 length (measured medially) 8.5; Gt 4 length (measured medially) 15; Gt 7 length (measured medially) 18; Gt 7 width (measured at base) 18; longest cercal seta length 19; shortest cercal seta length 7.

Male. Length of body 1.7–2.0mm.

Antenna ( Fig. 986) with scape widest just above the middle; plaque dark brown and situated in apical half, but distinctly behind apex; dorso-basal whorls of long setae present on F1–F4, with a ventro-basal whorl of long setae on C1. Gaster long ovate. Otherwise as in female.

Relative measurements: head length, dorsal view 24.5; head length, frontal view 39; head width 54; mouth width 20; malar space 15; eye length 25; scape length 20; scape width 7; plaque length 6; pedicel length 6; pedicel+flagellum length 92; F1 length 7; F1 width 6; F2 length 10.5; F2 width 6; F3 length 14; F3 width 5.5; F4 length 15; F4 width 4.5; clava length 36; clava width 4.5; mesosoma length 71; mesosoma width 48; gaster length 71; gaster width 34; longest subbasal seta on F1, length 47.

Hosts. Reared from leaf galls possibly induced by a Eurytomidae (though this requires confirmation) on Myrcia splendens ( Myrtaceae ). The gall is large (up to 2 cm in diameter) and spherical, sometimes transforming nearly the entire leaf into a gall; the interior consists numerous fibers in which are embedded several enclosed, whitish chambers, like seeds in a spongy matrix (P. Hanson, unpublished) ( Fig. 9).

Distribution. Costa Rica.

Material examined.

Holotype ♀ COSTA RICA, San José, Salitral de Santa Ana , 27.ii.1991, from Myrcia spongy leaf gall, P. Hanson ( NHMUK) . Paratypes (2♀ 1♂, MZLU, NHMUK) : 1♀ 1♂ with same label data as holotype ; 1♀ COSTA RICA, San José, Jerico, Desamparados , 1650m, v.1992, from gall on Myrcia splendens ”.

Etymology. Named after the host plant of the gall from which this species was reared.