Dima neumanni Platia, 2013

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 23-24

Dima neumanni Platia, 2013

Figs 67–74 , 146 , 199–201 , 233 , 246 .

Dima neumanni Platia, 2013: 98 .

Type depository. Holotype, ♂ (NME).

Type locality. Greece: Peloponnese, Mt. Erimanthos , SE Kalentzi, 1190 m.

Redetermined material. GREECE: nom. Ahaia, O. Aroania canalino, 1330 m, 1.VI.1995 –4. VI.1996, 1 ♀, Giachino & Vailati leg. (as D. hladilorum in Schimmel & Platia 2008 ) (PCGP).

New material. GREECE: distr. Ahaïa, Erímanthos Mts. , Kalentzi, firrywood, 1240 m (37°56'52.21"N, 21°46'15.90"E), 3. VI.2013, 1 ♂, 2 ♀♀, J. Mertlik leg. ( PCJM) ; dtto, 2 ♀♀, V. Dušánek leg. (PCVD); dtto, 2 ♀♀, P. Brůha leg. ( PCPB); distr. Ahaïa, Panachaiko Mts., Kalanistra , 20.IV.–2 . V.2014, 2 ♀♀, Manfred Egger leg. ( PCME); Erímanthos Mts., Kalentzi env. (ca. 37°56'49.41"N, 21°46'11.26"E), 21 . V.2015, 3 ♀♀, Manfred Egger leg. ( PCME); Erímanthos Mts., Kalentzi env., 1265 m (37°57'2.22"N, 21°46'35.46"E), 10 . VI.2015, 8 ♂♂, 2 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♀, B. Zbuzek leg. (PCBZ); dtto, 1 ♂, P. Brůha leg. (PCPB); dtto, 2 ♂♂, T. Németh leg. (PCRK); dtto, 1 ♀ ( HNHM); distr. Ahaïa, Panachaiko Mts., Kalanistra env., Agios Athanasios (ca. 38° 4'44.65"N, 21°51'35.45"E), 10 . V.2015, 2 ♂♂, 3 ♀♀, Manfred Egger leg. ( PCME); Peloponnese, Mt. Chelmos, Kalavrita env., 28 . IV.1993, 1 ♂, J. Macek leg. ( PCVK); distr. Ahaïa, Aroánia Óri Mts. , firrywood, 1420 m (38°1'14.26"N, 22°10'21.28"E), 30 . V.2013, 1 ♂, 1 ♀, J. Mertlik leg.; dtto, 30. V.2013, 1 ♂, P. Brůha leg. ( PCVD); Mt. Chelmos (38°0'54.31"N, 22°11'24.35"E), 12 . V.2015, 1 ♀, Manfred Egger leg. (PCME).

Description of female. Like male, but more robust body, shorter antennae, surpassing the posterior angles of pronotum about two segments (more than three in males), and pronotum wider ( Figs 69, 71–74 ). The sclerotized spines of bursa copulatrix as in Fig. 233 .

Diagnosis. Dima neumanni is a medium-sized to large species (body length: 11.5–14.7 mm) with the coarse pronotal punctation, the long straight erect setae on the pronotal sides ( Fig. 146 ), and long straight semi-erect to erect pubescence of elytra ( Figs 67–74 ). The species limits of D. neumanni and D. schnitteri are unclear and it is possible, that they are conspecific. Dima schnitteri is known only from the lower elevations of the southern part of the Erimanthos Mts. (south from the distributional range of D. neumanni ; Fig. 246 ), is mostly larger, more flattened and paler ( Figs 105–108 ). The third species in the region, D. assingi , is also morphologically very like D. neumanni , but differs in the elytra with shorter decumbent to semi-erect pubescence in basal third or half, and in not erect (only semi-erect) setae in the first halfs of the pronotal sides ( Figs 6–9 , 120 ).

Intraspecific variability. Dima neumanni is rather a morphologically variable species, with the long erect pubescence of elytra as the only stable character. The apical part of the scutellum is either more or less rounded or subacute. The specimens from Aroania Mts. are paler, and one of the males has more elongate apical lobe of paramera. All known specimens from Aroania Mts. and some from Panachaiko Mts. have more parallel-sided and elongate pronotum than the specimens from the type locality in Erimanthos Mts. ( Figs 67–74 ). It must be noted, that all specimens of D. assingi (which differs only slightly from D. neumanni and occurs also in Aroania Mts.) are lightly colored and have also more elongate pronotum ( Figs 6–9 ). The limits of the Dima species distributed in the northern Peloponnese must be confirmed by future studies including much more extensive sampling and the DNAbased methods of species identification.

Distribution. Greece (Peloponnese: Erimanthos Mts., Panachaiko Mts., Aroania Mts.; Fig. 246 ).

Figures

FIGURE 246. The distribution of Dima species in the Southern Greece.

FIGURES 60 – 74. Habitus images of Dima species. Dima macedonica Schimmel, 1993, 60 — non-type male from Ehloec, 61 — female paratype of D. hladilorum Schimmel, 1987 from Galičica Mts.; D. marvani Mertlik & Dusanek, 2006, 62 — holotype, male, 63 — female paratype from Lovčen Mts., 64 — female paratype from Rumija Mts., 65 — non-type male from Rumija Mts.; 66 — D. moraveci Schimmel & Platia 2008, holotype, female; D. neumanni Platia, 2013, 67 — holotype, male, 68 — non-type male from Erimanthos Mts., 69 — non-type female from Erimanthos Mts., 70 — non-type male from Aroania Mts., 71 — non-type female from Aroania Mts., 72 — non-type female from Aroania Mts., 73 — non-type female from Panachaiko Mts., 74 — non-type female from Panachaiko Mts. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 192 – 201. Aedeagus images of Dima species. 192 — D. lebenbaueri Schimmel & Platia 2008, holotype; 193 — D. lebenbaueri, non-type specimen; 194 — D. macedonica Schimmel, 1993, holotype; 195 — D. macedonica, paratype; 196 — D. macedonica, non-type specimen from Galičica Mts.; 197 — D. macedonica, non-type specimen from Ehloec; 198 — D. marvani Mertlik & Dusanek, 2006, holotype; 199 — D. neumanni Platia, 2013, holotype; 200 — D. neumanni, non-type specimen from Erimanthos Mts.; 201 — D. neumanni, non-type specimen from Aroania Mts. Not to scale.

FIGURES 222 – 233. The sclerotized spines of bursa copulatrix of Dima species. 222 — D. arndti Platia, 2013, paratype; 223 — D. bruhai sp. nov., paratype; 224 — D. dusaneki sp. nov., paratype; 225 — D. elateroides, Charpentier, 1825, non-type specimen; 226 — D. etoliensis Platia, 2012, holotype; 227 — D. fialai sp. nov., paratype; 228 — D. florinensis Platia, 2012, holotype; 229 — D. fthiotidensis Schimmel & Platia, 2008, holotype of D. scutellaris Platia, 2010; 230 — D. giachinoi Schimmel & Platia 2008, paratype; 231 — D. hirtipennis Platia, 2011, holotype; 232 — D. kozufensis sp. nov., paratype; 233 — D. neumanni Platia, 2013, non-type specimen. Not to scale.

FIGURES 105 – 118. Habitus images of Dima species. Dima schnitteri Platia, 2013, 105 — holotype, male, 106 — paratype, female, 107 — non-type specimen, male, 108 — non-type specimen, female; D. vailatii Schimmel & Platia 2008, 109 — holotype, male, 110 — paratype, female, 111 — non-type specimen, male, 112 — non-type specimen, female, 113 — holotype of D. riesei Platia, 2010, male; D. vonickai sp. nov., 114 — holotype, male, 115 — paratype, female; D. zbuzeki sp. nov., 116 — holotype, male, 117 — paratype, female, 118 — paratype, male. Not to scale.

FIGURES 1 – 14. Habitus images of Dima species. Dima arndti Platia, 2013, 1 — holotype, male, 2 — paratype, female, 3 — nontype specimen, male, 4 — non-type specimen, female; 5 — Dima sp. from Crete, male; D. assingi Schimmel & Platia, 2008, 6 — holotype, male, 7 — holotype of D. peloponnensis Schimmel & Platia, 2008, male, 8 — non-type male from Kilini Mts., 9 — nontype female from Kilini Mts.; D. bruhai sp. nov., 10 — holotype, male, 11 — paratype, female, 12 — paratype, male, 13 — paratype, male; D. dalmatina Küster, 1844, 14 — syntype, male, lateral view. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.